^1.Guangxi Key Laboratory of Plant Conservation and Restoration Ecology in Karst Terrain, Guangxi Institute of Botany, Chinese Academy of Sciences, Guilin 541006, China

^2.Guangxi Guilin Urban Ecosystem National Observation and Research Station, National Forestry and Grassland Administration, Guilin 541006, China

^3.State Key Laboratory of Desert and Oasis Ecology, Xinjiang Institute of Ecology and Geography, Chinese Academy of Sciences, Urumqi 830011, China

^4.Cele National Station of Observation and Research for Desert-Grassland Ecosystems, Cele 848300, China

^5.College of Tourism, Kaili University, Kaili 556011, China

^6.Guangdong Provincial Key Laboratory of Applied Botany, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 510650, China

^7.Guangdong Province Data Center of Terrestrial and Marine Ecosystems Carbon Cycle, Guangzhou 510650, China