^1.Key Laboratory of Biodiversity and Environment on the Qinghai-Tibetan Plateau, Ministry of Education, School of Ecology and Environment, Xizang University, Lhasa 850000, China

^2.State Key Laboratory of Severe Weather Meteorological Science and Technology, Chinese Academy of Meteorological Sciences, Beijing 100081, China

^3.Northwest Institute of Eco-Environment and Resources, Chinese Academy of Sciences, Lanzhou 730000, China

^4.Institute of Mountain Hazards and Environment, Chinese Academy of Sciences, Chengdu 610213, China

^5.State Key Laboratory of Tibetan Plateau Earth System, Environment and Resources, Institute of Tibetan Plateau Research Chinese Academy of Sciences, Beijing 100101, China