^1.a. Hunan Provincial Key Laboratory for Control of Forest Diseases and Pests; b. Key Laboratory of Cultivation and Protection for Non-wood Forest Trees, Ministry of Education; c. Key Laboratory of Integrated Management of Forest Biological Resources and Pests of Universities in Hunan Province; d. College of Life and Environmental Sciences, Central South University of Forestry & Technology, Changsha 410004, Hunan, China

^2.Guangxi State-owned Bobai Forest Farm, Yulin 537600, Guangxi, China

^3.a. Zhongnan Fast-growing Wood Breeding Laboratory of National Forestry and Grassland Administration; b. Guangxi Key Laboratory of Superior Timber Tree Resource Cultivation, Guangxi Forestry Research Institute, Nanning 530002, Guangxi, China

^1d.College of Life and Environmental Sciences, Central South University of Forestry & Technology, Changsha 410004, Hunan, China