¹College of Environmental and Resource Sciences, Zhejiang University, Hangzhou 310058, China

²Zhejiang Provincial Key Laboratory of Agricultural Resources and Environment, Zhejiang University, Hangzhou 310058, China

³School of Agriculture and Food, The University of Melbourne, Melbourne, VIC 3010, Australia

⁴Key Laboratory of the Three Gorges Reservoir Region’s Eco-Environment, Ministry of Education, Chongqing University, Chongqing 400045, China

⁵Department of Biology, Stanford University, Stanford, CA 94305, USA

⁶Key Lab of Urban Environment and Health, Institute of Urban Environment, Chinese Academy of Sciences, Xiamen, Fujian 361021, China

⁷Research Center for Eco-Environmental Sciences, Chinese Academy of Sciences, Beijing 100085, China