¹ State Key Laboratory of Coastal and Offshore Engineering, Dalian University of Technology, Dalian 116024, China.

² State Key Laboratory of Bioelectronics, School of Biological Science and Medical Engineering, Southeast University, Nanjing 210096, China.

³ State Key Laboratory of Marine Resources Utilization in South China Sea, Hainan University, Haikou 570228, China.

⁴ Guangxi Key Lab of Mangrove Conservation and Utilization, Guangxi Mangrove Research Center, Guangxi Academy of Sciences, Beihai 536000, China.

⁵ Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China.

⁶ Southern Marine Science and Engineering Guangdong Laboratory (Guangzhou), Guangzhou 511458, China.

⁷ South China Sea Institute of Oceanology, Chinese Academy of Science, Guangzhou 510301, China.