¹Department of Biomedical Engineering, Southern University of Science and Technology, Shenzhen 518055, Guangdong, China

²Department of Biochemistry and Molecular Biology, School of Basic Medical Sciences, Peking University, Beijing 100191, China

³Department of Structural Biology, St. Jude Children’s Research Hospital, Memphis, TN 38105, USA

⁴Center for Protein Science and Crystallography, School of Life Sciences, Faculty of Science, Chinese University of Hong Kong, Hong Kong 999077, China

⁵Interdisciplinary Center for Brain Information, the Brain Cognition and Brain Disease Institute, Shenzhen Institute of Advanced Technology, Chinese Academy of Sciences, Shenzhen 518055, Guangdong, China

⁶Faculty of Life and Health Sciences, Shenzhen Institute of Advanced Technology, Chinese Academy of Sciences, Shenzhen 518055, Guangdong, China

⁷Shenzhen-Hong Kong Institute of Brain Science-Shenzhen Fundamental Research Institutions, Shenzhen 518055, Guangdong, China

⁸Shenzhen Institute of Advanced Technology, Chinese Academy of Sciences, Shenzhen 518055, Guangdong, China

⁹State Key Lab for Biocontrol, School of Life Sciences, Sun Yat-Sen University, Guangzhou 510275, China

¹⁰Cryo-EM Facility Center, Southern University of Science and Technology, Shenzhen 518055, Guangdong, China

¹¹Department of Orthopedics, Shenzhen Intelligent Orthopaedics and Biomedical Innovation Platform, Guangdong Provincial Research Center for Artificial Intelligence and Digital Orthopedic Technology, Shenzhen Second People’s Hospital, the First Affiliated Hospital of Shenzhen University, Shenzhen 518000, Guangdong, China

¹²Guangdong Provincial Key Laboratory of Advanced Biomaterials, Southern University of Science and Technology, Shenzhen 518055, Guangdong, China