¹ Indian Ocean Ecosystem Research Center, Tianjin University of Science and Technology, Tianjin 300457, China

² Ministry of Education Key Laboratory of the Coastal and Wetland Ecosystems, College of the Environment Ecology, Xiamen University, Xiamen 361102, China

³ Key Laboratory of Marine Ecosystem and Biogeochemistry, Second Institute of Oceanography, Ministry of Natural Resources, Hangzhou 310012, China

⁴ Ministry of Agriculture Key Laboratory of Sustainable Development of Marine Fisheries, Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Qingdao 266071, China

⁵ Key Laboratory of Animal and Plant Resistance in Tianjin, College of Life Science of Tianjin Normal University, Tianjin 300387, China

⁶ Third Institute of Oceanography, Ministry of Natural Resources, Xiamen 361005, China

⁷ Institute of Marine Science and Technology, Shandong University, Qingdao 266237, China

⁸ Department of Marine Organism Taxonomy and Phylogeny, Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, China

⁹ College of Marine Life Sciences, Ocean University of China, Qingdao 266003, China