^a Xiamen Key Laboratory of Clean and High-valued Utilization for Biomass, College of Energy, Xiamen University, Xiamen 361102, China

^b Fujian Engineering and Research Center of Clean and High-valued Technologies for Biomass, Xiamen 361102, China

^c School of Chemical and Biological Engineering, Zhejiang University, Hangzhou 310058, China

^d Department of Paper and Bioprocess Engineering, State University of New York College of Environmental Science and Forestry, Syracuse, New York 13210, United States

^e State Key Laboratory of Applied Microbiology Southern China, Guangdong Provincial Key Laboratory of Microbial Culture Collection and Application, Guangdong Open Laboratory of Applied Microbiology, Guangdong Microbial Culture Collection Center (GDMCC), Guangdong Institute of Microbiology, Guangdong Academy of Sciences, Guangzhou 510070, China