1 Key Laboratory of Marine Ecosystem and Biogeochemistry, Second Institute of Oceanography, State Oceanic Administration, Hangzhou 310012, China
2 Laboratory for Marine Biology and Biotechnology, Pilot National Laboratory for Marine Science and Technology (Qingdao), Qingdao 266237, China
3 Center for Ocean Mega-Science, Chinese Academy of Sciences, Qingdao 266071, China
4 Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, China
5 University of Chinese Academy of Sciences, Beijing 100049, China
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