¹ National Observation and Research Station for the Taiwan Strait Marine Ecosystem/Fujian Provincial Key Laboratory of Coastal Ecology and Environmental Studies, College of the Environment and Ecology, Xiamen University, Xiamen 361102, China

² Fujian Province Key Laboratory for the Development of Bioactive Material from Marine Algae, College of Resources and Environmental Sciences, Quanzhou Normal University, Quanzhou 362000, China

³ Southern Marine Science and Engineering Guangdong Laboratory (Zhuhai), Zhuhai 511458, China

⁴ Department of Environmental Sciences, College of the Coast & Environment, Louisiana State University, Baton Rouge 70803, LA, USA