¹ College of Geography and Environmental Science, Key Laboratory of Tropical Island Land Surface Processes and Environmental Changes of Hainan Province, Hainan Normal University, Haikou 571158, China

² State Key Laboratory of Estuarine and Coastal Research, East China Normal University, Shanghai 200241, China

³ State Key Laboratory of Marine Resource Utilization in South China Sea, Hainan University, Haikou 570228, China

⁴ Shanghai Key Laboratory of Polar Life and Environment Sciences, School of Oceanography, Shanghai Jiao Tong University, Shanghai 200030, China

⁵ MNR Key Laboratory for Polar Science, Polar Research Institute of China, Shanghai 200136, China

⁶ Key Laboratory of Coastal Environment and Resources of Zhejiang Province, School of Engineering, Westlake University, Hangzhou 310024, China