^1.Spice and Beverage Research Institute, Chinese Academy of Tropical Agricultural Science, Wanning 571533, China

^2.Key Laboratory of Genetic Resources Utilization of Spice and Beverage Crops, Ministry of Agriculture and Rural Affairs, Wanning 571533, China

^3.Hainan Provincial Key Laboratory of Genetic Improvement and Quality Regulation for Tropical Spice and Beverage Crops, Wanning 571533, China

^4.Tropical Crops College, Yunnan Agricultural University, Puer 665000, China