Article(id=1305135917799330409, tenantId=1146029695717560320, journalId=1303687221787852879, issueId=1305135915303723476, articleNumber=null, orderNo=null, doi=10.7606/j.issn.1009-1041.2026.07.02, pmid=null, cstr=null, oa=null, hot=null, price=null, onlineType=0, articleFormat=0, articleType=null, articleTypeStr=null, receivedDate=1768060800000, receivedDateStr=2026-01-11, revisedDate=1773763200000, revisedDateStr=2026-03-18, acceptedDate=null, acceptedDateStr=null, onlineDate=1789098251055, onlineDateStr=2026-09-11, pubDate=1784044800000, pubDateStr=2026-07-15, doiRegisterDate=null, doiRegisterDateStr=null, onlineIssueDate=1789098251055, onlineIssueDateStr=2026-09-11, onlineJustAcceptDate=null, onlineJustAcceptDateStr=null, onlineFirstDate=null, onlineFirstDateStr=null, sourceXml=null, magXml=null, createTime=1789098251055, creator=13701087609, updateTime=1789098251055, updator=13701087609, issue=Issue{id=1305135915303723476, tenantId=1146029695717560320, journalId=1303687221787852879, year='2026', volume='46', issue='7', pageStart='F2', pageEnd='990', issueExtLink='null', onlineDate='null', pubDate='1784044800000', pubDateStr='2026-07-15', beforeIssueId=null, nextIssueId=null, price=null, status=1, issueComplete=1, articleOrder=1, issueType=-1, specialIssue=null, createTime=1789098250461, creator='13701087609', updateTime=1789103040607, updator='13701087609', preIssue=null, nextIssue=null, articleTotal=null, ext={EN=IssueExt(id=1305156006661608318, tenantId=1146029695717560320, journalId=1303687221787852879, issueId=1305135915303723476, language=EN, specialIssueTitle=, coverIllustrator=null, specialIssueEditor=, specialIssueAbout=), CN=IssueExt(id=1305156006661608319, tenantId=1146029695717560320, journalId=1303687221787852879, issueId=1305135915303723476, language=CN, specialIssueTitle=, coverIllustrator=null, specialIssueEditor=, specialIssueAbout=)}, issueFiles=null, downloadFileDto=null}, startPage=862, endPage=870, ext={EN=ArticleExt(id=1305135918021628522, articleId=1305135917799330409, tenantId=1146029695717560320, journalId=1303687221787852879, language=EN, title=Effects of TaNAK1.1 and TaNAK1.2 on Agronomic and Yield-Related Traits and Drought Tolerance in Wheat, columnId=1305133119129023428, journalTitle=Journal of Triticeae Crops, columnName=Genetics & Breeding, runingTitle=null, highlight=null, articleAbstract=

To investigate the effects of two alternative splicing variants of wheat receptor-like kinase gene TaNAK1 on agronomic and yield-related traits, this study used the winter wheat cultivar Xiaoyan 6, the wild-type spring wheat cultivar Fielder, and its transgenic lines overexpressing TaNAK1.1 (OE1-1, OE1-9, OE1-11） and TaNAK1.2 (OE2-1, OE2-2, OE2-4). The expression patterns of TaNAK1.1 and TaNAK1.2 in wheat were examined using semi-quantitative RT-PCR (sqRT-PCR) and quantitative real-time RT-PCR (qRT-PCR), respectively. Agronomic and yield-related traits of the transgenic lines under field conditions, along with seedling drought tolerance, were evaluated. The results showed that TaNAK1.1 and TaNAK1.2 exhibited distinct spatiotemporal expression profiles across wheat organs and tissues under normal conditions; TaNAK1.1 displayed relatively high expression levels in grains at 10 and 15 days post-anthesis, whereas TaNAK1.2 showed relatively high expression levels in seedling leaves and flag leaves. Under drought and low-temperature stress conditions, TaNAK1.2 was significantly up-regulated, while the expression of TaNAK1.1 was not induced by these abiotic stresses. The overexpression of TaNAK1.1 and TaNAK1.2 differentially affected agronomic and yield-related traits under field conditions. Compared to wild-type wheat Fielder, TaNAK1.1 transgenic lines showed no significant differences in heading and flowering time or flag leaf area. In contrast, heading and flowering time in TaNAK1.2 transgenic lines were delayed by approximately 5 d, and flag leaf area was reduced by 12.50% to 14.45%. Both TaNAK1.1 transgenic and TaNAK1.2 lines exhibited significant reductions in plant height, tiller number, spike length, number of spikelets per spike, grain size, thousand-grain weight, and grain weight per plant; however, the extent of these reductions varied between the two types of overexpressing lines, with TaNAK1.1 transgenic lines showing greater reductions in grain weight per plant and biomass per plant. The effects of TaNAK1.1 and TaNAK1.2 overexpression on wheat seedling drought tolerance were similar, with both significantly enhancing drought tolerance. Specifically, under drought stress, compared to the wild-type wheat Fielder, the leaves of TaNAK1.1 transgenic and TaNAK1.2 transgenic lines showed significantly increased contents of osmotic adjustment solutes (soluble sugars and proline) and relative water content, along with significantly decreased levels of H₂O₂ and the membrane lipid peroxidation product MDA. Furthermore, their survival rate was also significantly enhanced following extreme drought stress treatment. These results not only reveal the important roles of TaNAK1 alternative splicing in regulating growth, development, and adaptation to abiotic stress in wheat, but also provide new genetic resources for molecular breeding of stress-tolerant wheat.

, authors=Yue GONG, Kunzhi HU, Baowei WU, Didi DU, Meng MA, Xiangli LIU, Huixian ZHAO, authorsList=Yue GONG, Kunzhi HU, Baowei WU, Didi DU, Meng MA, Xiangli LIU, Huixian ZHAO, authorCompany=null, correspAuthors=Huixian ZHAO, authorNote=null, correspAuthorsNote=null, copyrightStatement=null, copyrightOwner=null, extLink=null, articleAbsUrl=null, sourceXml=null, magXml=null, pdfUrl=null, pdf=null, pdfFileSize=null, pdfExtLink=null, richHtmlUrl=null, mobilePdfUrl=null, reviewReport=null, pdfFirstPage=null, abstractGraph=null, abstractGraphContent=null, abstractVideo=null, citation=null, cebUrl=null, magXmlContent=null, mapNumber=null, fund=null), CN=ArticleExt(id=1305135918403310189, articleId=1305135917799330409, tenantId=1146029695717560320, journalId=1303687221787852879, language=CN, title=TaNAK1.1和TaNAK1.2对小麦农艺和产量相关性状及耐旱性的效应, columnId=1305133119284212678, journalTitle=麦类作物学报, columnName=遗传育种, runingTitle=null, highlight=null, articleAbstract=

为探究小麦类受体激酶基因TaNAK1的两个可变剪接体对小麦农艺和产量相关性状的影响，以冬小麦品种小偃6号、野生型春小麦品种Fielder及其TaNAK1.1（OE1-1、OE1-9、OE1-11）和TaNAK1.2转基因株系（OE2-1、OE2-2、OE2-4）为材料，采用半定量RT-PCR和实时荧光定量RT-qPCR分别检测TaNAK1.1和TaNAK1.2在小麦中的表达模式，并对转基因小麦株系的田间农艺和产量相关性状及幼苗耐旱性进行评价。结果表明，小麦TaNAK1.1和TaNAK1.2在正常条件下的时空表达谱及其在干旱和低温处理下的表达模式均存在差异。正常条件下，TaNAK1.1在花后10 d和15 d籽粒中表达水平相对较高，而TaNAK1.2在幼苗叶片和旗叶中表达水平较高；干旱和低温处理条件下，TaNAK1.2的表达显著上调，而TaNAK1.1的表达变化不明显。TaNAK1.1和TaNAK1.2过表达对小麦田间农艺和产量相关性状的影响不同。与野生型Fielder相比，TaNAK1.1转基因株系的抽穗、扬花时间和旗叶面积差异不大；而TaNAK1.2转基因株系的抽穗和扬花时间均延后约5 d，旗叶面积减小12.50%~14.45%；TaNAK1.1和TaNAK1.2转基因株系的株高、分蘖数、穗长、主穗小穗数、籽粒大小、千粒重、单株产量等性状均显著降低，但二者间各性状降低幅度不尽相同，TaNAK1.1转基因株系的单株产量和单株生物量的降幅更大。TaNAK1.1和TaNAK1.2过表达均能显著提升小麦的耐旱性。在干旱胁迫下，与野生型Fielder相比，TaNAK1.1和TaNAK1.2转基因株系叶片相对含水量、可溶性糖和脯氨酸含量均显著增加，H₂O₂和MDA含量均显著降低；经极度干旱胁迫处理后，转TaNAK1.1和TaNAK1.2株系的存活率均显著提高。综上所述，TaNAK1.1和TaNAK1.2在调控小麦生长发育与抗非生物胁迫中有重要作用，可为小麦抗逆分子育种提供新的基因资源。

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E-mail: gongyue0214@nwafu.edu.cn（龚悦）

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E-mail: gongyue0214@nwafu.edu.cn（龚悦）

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