Article(id=1305139346928071460, tenantId=1146029695717560320, journalId=1303687221787852879, issueId=1305139324555649224, articleNumber=null, orderNo=null, doi=10.7606/j.issn.1009-1041.2026.03.04, pmid=null, cstr=null, oa=null, hot=null, price=null, onlineType=0, articleFormat=0, articleType=null, articleTypeStr=null, receivedDate=1743523200000, receivedDateStr=2025-04-02, revisedDate=1747756800000, revisedDateStr=2025-05-21, acceptedDate=null, acceptedDateStr=null, onlineDate=1789099068623, onlineDateStr=2026-09-11, pubDate=1773504000000, pubDateStr=2026-03-15, doiRegisterDate=null, doiRegisterDateStr=null, onlineIssueDate=1789099068623, onlineIssueDateStr=2026-09-11, onlineJustAcceptDate=null, onlineJustAcceptDateStr=null, onlineFirstDate=null, onlineFirstDateStr=null, sourceXml=null, magXml=null, createTime=1789099068623, creator=13701087609, updateTime=1789099068623, updator=13701087609, issue=Issue{id=1305139324555649224, tenantId=1146029695717560320, journalId=1303687221787852879, year='2026', volume='46', issue='3', pageStart='F2', pageEnd='424', issueExtLink='null', onlineDate='null', pubDate='1773504000000', pubDateStr='2026-03-15', beforeIssueId=null, nextIssueId=null, price=null, status=1, issueComplete=1, articleOrder=1, issueType=-1, specialIssue=null, createTime=1789099063289, creator='13701087609', updateTime=1789103147683, updator='13701087609', preIssue=null, nextIssue=null, articleTotal=null, ext={EN=IssueExt(id=1305156455804465602, tenantId=1146029695717560320, journalId=1303687221787852879, issueId=1305139324555649224, language=EN, specialIssueTitle=, coverIllustrator=null, specialIssueEditor=, specialIssueAbout=), CN=IssueExt(id=1305156455804465603, tenantId=1146029695717560320, journalId=1303687221787852879, issueId=1305139324555649224, language=CN, specialIssueTitle=, coverIllustrator=null, specialIssueEditor=, specialIssueAbout=)}, issueFiles=null, downloadFileDto=null}, startPage=312, endPage=320, ext={EN=ArticleExt(id=1305139347137786661, articleId=1305139346928071460, tenantId=1146029695717560320, journalId=1303687221787852879, language=EN, title=Identification and Expression Characteristics Analysis of the Grain Hardness Gene HvHinb-1 in Hulless Barley, columnId=1305133119129023428, journalTitle=Journal of Triticeae Crops, columnName=Genetics & Breeding, runingTitle=null, highlight=null, articleAbstract=

To obtain the superior genotype of HvHinb-1 gene related to the hardness of hulless barley grains, two hulless barley varieties with significant differences in hardness were seleeted: Suma (hardness value: 14.66±1.55) and Nanmulin (hardness value: 21.42±1.83). The HvHinb-1 gene sequence through PCR were doned to analyzed the sequence characteristics, protein physicochemical properties, and expression patterns of different genotypes, and explored their roles in the formation of hulless barley grain hardness. The distribution and genotype of HvHinb-1 in 21 hulless barley varieties with different hardness were analyzed, and the expression of HvHinb-1 in different stages of grain development (early, middle and late milk ripening) was detected by real-time quantitative PCR (qRT-PCR).The results showed that the HvHinb-1 was cloned from both the Suma and Nanmulin, with a sequence identity of 98.64%, and two amino acid differences. HvHinb-1 is a hydrophilic basic protein with an isoelectric point of 8.69 and a molecular weight of 16.12 to 16.14 kDa, primarily composed of α-helices and random coils. Phylogenetic analysis indicated that hulless barley HvHinb-1 has the closest genetic relationship to wheat. In the preliminary identification of 12 samples of low-hardness hulless barley (hardness value range: 9.04-15.04), the amino acid sequence at the mutation sites 78 and 94 was H, and in 9 samples of high-hardness hulless barley (hardness value range: 18.93-22.76), the amino acid sequence at the mutation sites 78 and 94 was Q. qRT-PCR analysis showed that in early milk ripening grains, the expression level of HvHinb-1 was significantly higher in Suma than that in Nanmulin, and in mid-milk ripening, it was extremely higher in Nanmulin than in Suma, suggesting that this period may be crucial for grain hardness formation.

, authors=Fenggui GUO^{1, 3}, Yating PENG^{1, 3}, Yan WANG^{2, 3}, Rundong XIE^{2, 3}, Youhua YAO^{2, 3}, Xiaohua YAO^{2, 3}, authorsList=Fenggui GUO, Yating PENG, Yan WANG, Rundong XIE, Youhua YAO, Xiaohua YAO, authorCompany=null, correspAuthors=Xiaohua YAO, authorNote=null, correspAuthorsNote=null, copyrightStatement=null, copyrightOwner=null, extLink=null, articleAbsUrl=null, sourceXml=null, magXml=null, pdfUrl=null, pdf=null, pdfFileSize=null, pdfExtLink=null, richHtmlUrl=null, mobilePdfUrl=null, reviewReport=null, pdfFirstPage=null, abstractGraph=null, abstractGraphContent=null, abstractVideo=null, citation=null, cebUrl=null, magXmlContent=null, mapNumber=null, fund=null), CN=ArticleExt(id=1305139347481719590, articleId=1305139346928071460, tenantId=1146029695717560320, journalId=1303687221787852879, language=CN, title=青稞籽粒硬度基因HvHinb-1鉴定及其表达特性分析, columnId=1305133119284212678, journalTitle=麦类作物学报, columnName=遗传育种, runingTitle=null, highlight=null, articleAbstract=

为获得青稞籽粒硬度相关基因HvHinb-1的优异基因型，选取硬度差异显著的青稞品种苏马（硬度值：14.66±1.55）和南木林（硬度值：21.42±1.83），通过PCR克隆并测序获得HvHinb-1基因序列，分析不同基因型序列特征、蛋白理化性质及表达模式；分析HvHinb-1在21份硬度不同的青稞品种的分布及基因型，并利用实时荧光定量PCR（qRT-PCR）技术检测HvHinb-1在籽粒发育不同阶段（乳熟早期、中期和晚期）的表达量。结果表明，从青稞品种苏马和南木林中均可克隆到HvHinb-1基因，其编码蛋白序列一致性为98.64%，存在2个氨基酸差异。HvHinb-1为亲水性碱性蛋白，等电点8.69，分子量16.12~16.14 kDa，二级结构组成以α-螺旋和无规卷曲为主。经系统进化分析，青稞与大麦HvHinb-1亲缘关系最近。在12份硬度较小（硬度：9.04~15.04）青稞材料中，HvHinb-1的氨基酸序列在第78和94处为H，9份硬度较大（硬度：18.93~22.76）青稞材料中，其氨基酸序列在第78和94处为Q。通过qRT-PCR分析，在乳熟早期籽粒中，HvHinb-1在苏马中的表达量显著高于南木林，而在乳熟中期，在南木林中表达量极显著高于苏马，说明该时期可能是籽粒硬度形成的关键阶段。

, authors=郭凤贵^{1, 3}, 彭雅婷^{1, 3}, 王燕^{2, 3}, 谢润东^{2, 3}, 姚有华^{2, 3}, 姚晓华^{2, 3}, authorsList=郭凤贵, 彭雅婷, 王燕, 谢润东, 姚有华, 姚晓华, authorCompany=null, correspAuthors=姚晓华, authorNote=, correspAuthorsNote=, copyrightStatement=null, copyrightOwner=null, extLink=null, articleAbsUrl=null, sourceXml=AsvydtoP1IvprRiu9F1G3w==, magXml=42mEquyX1X6b8LCeEGC56g==, pdfUrl=null, pdf=pJtA0IySGJaSVwJKsQnVrQ==, pdfFileSize=5941828, pdfExtLink=null, richHtmlUrl=null, mobilePdfUrl=null, reviewReport=null, pdfFirstPage=null, abstractGraph=null, abstractGraphContent=null, abstractVideo=null, citation=null, cebUrl=null, magXmlContent=nx6guvQUBmg0fqbPtsXqdQ==, mapNumber=null, fund=null)}, authors=[Author(id=1305139348228305713, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, orderNo=0, firstName=null, middleName=null, lastName=null, nameCn=null, orcid=null, stid=null, country=null, authorPic=null, dead=0, email=2790066752@qq.com, emailSecond=null, emailThird=null, correspondingAuthor=0, authorType=1, ext={EN=AuthorExt(id=1305139348324774708, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, authorId=1305139348228305713, language=EN, stringName=Fenggui GUO, firstName=Fenggui, middleName=null, lastName=GUO, prefix=null, suffix=null, authorComment=null, nameInitials=null, affiliation=null, department=null, xref=^{1, 3}, address=^1.College of Agriculture and Animal Husbandry of Qinghai University, Xining, Qinghai 810016, China
^3.Qinghai Provincial Key Laboratory of Qingke Genetic Breeding/Qinghai Barley Subcenter of National Wheat Improvement Center/Laboratory of Germplasm Resources Research and Utilization on the Qinghai-Tibet Plateau, Xining, Qinghai 810016, China, bio=null, bioImg=null, bioContent=null, aboutCorrespAuthor=null), CN=AuthorExt(id=1305139348400272181, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, authorId=1305139348228305713, language=CN, stringName=郭凤贵, firstName=null, middleName=null, lastName=null, prefix=null, suffix=null, authorComment=null, nameInitials=null, affiliation=null, department=null, xref=^{1, 3}, address=^1．青海大学农牧学院，青海 西宁 810016
^3．青海省青稞遗传育种重点实验室/国家麦类改良中心青海青稞分中心/青藏高原种质资源研究与利用实验室，青海 西宁 810016, bio={"content":"

E-mail: 2790066752@qq.com（郭凤贵）

"}, bioImg=null, bioContent=

E-mail: 2790066752@qq.com（郭凤贵）

, aboutCorrespAuthor=null)}, companyList=[AuthorCompany(id=1305139347834041127, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, xref=1., ext=[AuthorCompanyExt(id=1305139347838235432, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, companyId=1305139347834041127, language=EN, country=null, province=null, city=null, postcode=null, companyName=null, departmentName=null, remark=^1.College of Agriculture and Animal Husbandry of Qinghai University, Xining, Qinghai 810016, China), AuthorCompanyExt(id=1305139347846624041, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, companyId=1305139347834041127, language=CN, country=null, province=null, city=null, postcode=null, companyName=null, departmentName=null, remark=^1．青海大学农牧学院，青海 西宁 810016)]), AuthorCompany(id=1305139348018590509, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, xref=3., ext=[AuthorCompanyExt(id=1305139348144419630, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, companyId=1305139348018590509, language=EN, country=null, province=null, city=null, postcode=null, companyName=null, departmentName=null, remark=^3.Qinghai Provincial Key Laboratory of Qingke Genetic Breeding/Qinghai Barley Subcenter of National Wheat Improvement Center/Laboratory of Germplasm Resources Research and Utilization on the Qinghai-Tibet Plateau, Xining, Qinghai 810016, China), AuthorCompanyExt(id=1305139348157002543, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, companyId=1305139348018590509, language=CN, country=null, province=null, city=null, postcode=null, companyName=null, departmentName=null, remark=^3．青海省青稞遗传育种重点实验室/国家麦类改良中心青海青稞分中心/青藏高原种质资源研究与利用实验室，青海 西宁 810016)])]), Author(id=1305139348479963959, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, orderNo=1, firstName=null, middleName=null, lastName=null, nameCn=null, orcid=null, stid=null, country=null, authorPic=null, dead=0, email=3508597744@qq.com, emailSecond=null, emailThird=null, correspondingAuthor=0, authorType=1, ext={EN=AuthorExt(id=1305139348651930426, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, authorId=1305139348479963959, language=EN, stringName=Yating PENG, firstName=Yating, middleName=null, lastName=PENG, prefix=null, suffix=null, authorComment=null, nameInitials=null, affiliation=null, department=null, xref=^{1, 3}, address=^1.College of Agriculture and Animal Husbandry of Qinghai University, Xining, Qinghai 810016, China
^3.Qinghai Provincial Key Laboratory of Qingke Genetic Breeding/Qinghai Barley Subcenter of National Wheat Improvement Center/Laboratory of Germplasm Resources Research and Utilization on the Qinghai-Tibet Plateau, Xining, Qinghai 810016, China, bio=null, bioImg=null, bioContent=null, aboutCorrespAuthor=null), CN=AuthorExt(id=1305139348719039291, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, authorId=1305139348479963959, language=CN, stringName=彭雅婷, firstName=null, middleName=null, lastName=null, prefix=null, suffix=null, authorComment=null, nameInitials=null, affiliation=null, department=null, xref=^{1, 3}, address=^1．青海大学农牧学院，青海 西宁 810016
^3．青海省青稞遗传育种重点实验室/国家麦类改良中心青海青稞分中心/青藏高原种质资源研究与利用实验室，青海 西宁 810016, bio={"content":"

3508597744@qq.com（彭雅婷）

"}, bioImg=null, bioContent=

3508597744@qq.com（彭雅婷）

, aboutCorrespAuthor=null)}, companyList=[AuthorCompany(id=1305139347834041127, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, xref=1., ext=[AuthorCompanyExt(id=1305139347838235432, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, companyId=1305139347834041127, language=EN, country=null, province=null, city=null, postcode=null, companyName=null, departmentName=null, remark=^1.College of Agriculture and Animal Husbandry of Qinghai University, Xining, Qinghai 810016, China), AuthorCompanyExt(id=1305139347846624041, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, companyId=1305139347834041127, language=CN, country=null, province=null, city=null, postcode=null, companyName=null, departmentName=null, remark=^1．青海大学农牧学院，青海 西宁 810016)]), AuthorCompany(id=1305139348018590509, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, xref=3., ext=[AuthorCompanyExt(id=1305139348144419630, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, companyId=1305139348018590509, language=EN, country=null, province=null, city=null, postcode=null, companyName=null, departmentName=null, remark=^3.Qinghai Provincial Key Laboratory of Qingke Genetic Breeding/Qinghai Barley Subcenter of National Wheat Improvement Center/Laboratory of Germplasm Resources Research and Utilization on the Qinghai-Tibet Plateau, Xining, Qinghai 810016, China), AuthorCompanyExt(id=1305139348157002543, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, companyId=1305139348018590509, language=CN, country=null, province=null, city=null, postcode=null, companyName=null, departmentName=null, remark=^3．青海省青稞遗传育种重点实验室/国家麦类改良中心青海青稞分中心/青藏高原种质资源研究与利用实验室，青海 西宁 810016)])]), Author(id=1305139348807119677, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, orderNo=2, firstName=null, middleName=null, lastName=null, nameCn=null, orcid=null, stid=null, country=null, authorPic=null, dead=0, email=null, emailSecond=null, emailThird=null, correspondingAuthor=0, authorType=1, ext={EN=AuthorExt(id=1305139348991669056, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, authorId=1305139348807119677, language=EN, stringName=Yan WANG, firstName=Yan, middleName=null, lastName=WANG, prefix=null, suffix=null, authorComment=null, nameInitials=null, affiliation=null, department=null, xref=^{2, 3}, address=^2.Qinghai University Academy of Agriculture and Forestry Sciences, Xining, Qinghai 810016, China
^3.Qinghai Provincial Key Laboratory of Qingke Genetic Breeding/Qinghai Barley Subcenter of National Wheat Improvement Center/Laboratory of Germplasm Resources Research and Utilization on the Qinghai-Tibet Plateau, Xining, Qinghai 810016, China, bio=null, bioImg=null, bioContent=null, aboutCorrespAuthor=null), CN=AuthorExt(id=1305139350208017217, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, authorId=1305139348807119677, language=CN, stringName=王燕, firstName=null, middleName=null, lastName=null, prefix=null, suffix=null, authorComment=null, nameInitials=null, affiliation=null, department=null, xref=^{2, 3}, address=^2．青海大学农林科学院，青海 西宁 810016
^3．青海省青稞遗传育种重点实验室/国家麦类改良中心青海青稞分中心/青藏高原种质资源研究与利用实验室，青海 西宁 810016, bio=null, bioImg=null, bioContent=null, aboutCorrespAuthor=null)}, companyList=[AuthorCompany(id=1305139347938898730, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, xref=2., ext=[AuthorCompanyExt(id=1305139347943093035, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, companyId=1305139347938898730, language=EN, country=null, province=null, city=null, postcode=null, companyName=null, departmentName=null, remark=^2.Qinghai University Academy of Agriculture and Forestry Sciences, Xining, Qinghai 810016, China), AuthorCompanyExt(id=1305139347951481644, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, companyId=1305139347938898730, language=CN, country=null, province=null, city=null, postcode=null, companyName=null, departmentName=null, remark=^2．青海大学农林科学院，青海 西宁 810016)]), AuthorCompany(id=1305139348018590509, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, xref=3., ext=[AuthorCompanyExt(id=1305139348144419630, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, companyId=1305139348018590509, language=EN, country=null, province=null, city=null, postcode=null, companyName=null, departmentName=null, remark=^3.Qinghai Provincial Key Laboratory of Qingke Genetic Breeding/Qinghai Barley Subcenter of National Wheat Improvement Center/Laboratory of Germplasm Resources Research and Utilization on the Qinghai-Tibet Plateau, Xining, Qinghai 810016, China), AuthorCompanyExt(id=1305139348157002543, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, companyId=1305139348018590509, language=CN, country=null, province=null, city=null, postcode=null, companyName=null, departmentName=null, remark=^3．青海省青稞遗传育种重点实验室/国家麦类改良中心青海青稞分中心/青藏高原种质资源研究与利用实验室，青海 西宁 810016)])]), Author(id=1305139350287708995, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, orderNo=3, firstName=null, middleName=null, lastName=null, nameCn=null, orcid=null, stid=null, country=null, authorPic=null, dead=0, email=null, emailSecond=null, emailThird=null, correspondingAuthor=0, authorType=1, ext={EN=AuthorExt(id=1305139350375789382, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, authorId=1305139350287708995, language=EN, stringName=Rundong XIE, firstName=Rundong, middleName=null, lastName=XIE, prefix=null, suffix=null, authorComment=null, nameInitials=null, affiliation=null, department=null, xref=^{2, 3}, address=^2.Qinghai University Academy of Agriculture and Forestry Sciences, Xining, Qinghai 810016, China
^3.Qinghai Provincial Key Laboratory of Qingke Genetic Breeding/Qinghai Barley Subcenter of National Wheat Improvement Center/Laboratory of Germplasm Resources Research and Utilization on the Qinghai-Tibet Plateau, Xining, Qinghai 810016, China, bio=null, bioImg=null, bioContent=null, aboutCorrespAuthor=null), CN=AuthorExt(id=1305139350438703943, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, authorId=1305139350287708995, language=CN, stringName=谢润东, firstName=null, middleName=null, lastName=null, prefix=null, suffix=null, authorComment=null, nameInitials=null, affiliation=null, department=null, xref=^{2, 3}, address=^2．青海大学农林科学院，青海 西宁 810016
^3．青海省青稞遗传育种重点实验室/国家麦类改良中心青海青稞分中心/青藏高原种质资源研究与利用实验室，青海 西宁 810016, bio=null, bioImg=null, bioContent=null, aboutCorrespAuthor=null)}, companyList=[AuthorCompany(id=1305139347938898730, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, xref=2., ext=[AuthorCompanyExt(id=1305139347943093035, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, companyId=1305139347938898730, language=EN, country=null, province=null, city=null, postcode=null, companyName=null, departmentName=null, remark=^2.Qinghai University Academy of Agriculture and Forestry Sciences, Xining, Qinghai 810016, China), AuthorCompanyExt(id=1305139347951481644, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, companyId=1305139347938898730, language=CN, country=null, province=null, city=null, postcode=null, companyName=null, departmentName=null, remark=^2．青海大学农林科学院，青海 西宁 810016)]), AuthorCompany(id=1305139348018590509, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, xref=3., ext=[AuthorCompanyExt(id=1305139348144419630, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, companyId=1305139348018590509, language=EN, country=null, province=null, city=null, postcode=null, companyName=null, departmentName=null, remark=^3.Qinghai Provincial Key Laboratory of Qingke Genetic Breeding/Qinghai Barley Subcenter of National Wheat Improvement Center/Laboratory of Germplasm Resources Research and Utilization on the Qinghai-Tibet Plateau, Xining, Qinghai 810016, China), AuthorCompanyExt(id=1305139348157002543, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, companyId=1305139348018590509, language=CN, country=null, province=null, city=null, postcode=null, companyName=null, departmentName=null, remark=^3．青海省青稞遗传育种重点实验室/国家麦类改良中心青海青稞分中心/青藏高原种质资源研究与利用实验室，青海 西宁 810016)])]), Author(id=1305139350530978633, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, orderNo=4, firstName=null, middleName=null, lastName=null, nameCn=null, orcid=null, stid=null, country=null, authorPic=null, dead=0, email=null, emailSecond=null, emailThird=null, correspondingAuthor=0, authorType=1, ext={EN=AuthorExt(id=1305139350610670412, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, authorId=1305139350530978633, language=EN, stringName=Youhua YAO, firstName=Youhua, middleName=null, lastName=YAO, prefix=null, suffix=null, authorComment=null, nameInitials=null, affiliation=null, department=null, xref=^{2, 3}, address=^2.Qinghai University Academy of Agriculture and Forestry Sciences, Xining, Qinghai 810016, China
^3.Qinghai Provincial Key Laboratory of Qingke Genetic Breeding/Qinghai Barley Subcenter of National Wheat Improvement Center/Laboratory of Germplasm Resources Research and Utilization on the Qinghai-Tibet Plateau, Xining, Qinghai 810016, China, bio=null, bioImg=null, bioContent=null, aboutCorrespAuthor=null), CN=AuthorExt(id=1305139350681973581, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, authorId=1305139350530978633, language=CN, stringName=姚有华, firstName=null, middleName=null, lastName=null, prefix=null, suffix=null, authorComment=null, nameInitials=null, affiliation=null, department=null, xref=^{2, 3}, address=^2．青海大学农林科学院，青海 西宁 810016
^3．青海省青稞遗传育种重点实验室/国家麦类改良中心青海青稞分中心/青藏高原种质资源研究与利用实验室，青海 西宁 810016, bio=null, bioImg=null, bioContent=null, aboutCorrespAuthor=null)}, companyList=[AuthorCompany(id=1305139347938898730, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, xref=2., ext=[AuthorCompanyExt(id=1305139347943093035, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, companyId=1305139347938898730, language=EN, country=null, province=null, city=null, postcode=null, companyName=null, departmentName=null, remark=^2.Qinghai University Academy of Agriculture and Forestry Sciences, Xining, Qinghai 810016, China), AuthorCompanyExt(id=1305139347951481644, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, companyId=1305139347938898730, language=CN, country=null, province=null, city=null, postcode=null, companyName=null, departmentName=null, remark=^2．青海大学农林科学院，青海 西宁 810016)]), AuthorCompany(id=1305139348018590509, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, xref=3., ext=[AuthorCompanyExt(id=1305139348144419630, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, companyId=1305139348018590509, language=EN, country=null, province=null, city=null, postcode=null, companyName=null, departmentName=null, remark=^3.Qinghai Provincial Key Laboratory of Qingke Genetic Breeding/Qinghai Barley Subcenter of National Wheat Improvement Center/Laboratory of Germplasm Resources Research and Utilization on the Qinghai-Tibet Plateau, Xining, Qinghai 810016, China), AuthorCompanyExt(id=1305139348157002543, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, companyId=1305139348018590509, language=CN, country=null, province=null, city=null, postcode=null, companyName=null, departmentName=null, remark=^3．青海省青稞遗传育种重点实验室/国家麦类改良中心青海青稞分中心/青藏高原种质资源研究与利用实验室，青海 西宁 810016)])]), Author(id=1305139350757471055, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, orderNo=5, firstName=null, middleName=null, lastName=null, nameCn=null, orcid=null, stid=null, country=null, authorPic=null, dead=0, email=yaoxiaohua009@126.com, emailSecond=null, emailThird=null, correspondingAuthor=1, authorType=1, ext={EN=AuthorExt(id=1305139350841357138, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, authorId=1305139350757471055, language=EN, stringName=Xiaohua YAO, firstName=Xiaohua, middleName=null, lastName=YAO, prefix=null, suffix=null, authorComment=null, nameInitials=null, affiliation=null, department=null, xref=^{2, 3}, address=^2.Qinghai University Academy of Agriculture and Forestry Sciences, Xining, Qinghai 810016, China
^3.Qinghai Provincial Key Laboratory of Qingke Genetic Breeding/Qinghai Barley Subcenter of National Wheat Improvement Center/Laboratory of Germplasm Resources Research and Utilization on the Qinghai-Tibet Plateau, Xining, Qinghai 810016, China, bio=null, bioImg=null, bioContent=null, aboutCorrespAuthor=null), CN=AuthorExt(id=1305139350921048915, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, authorId=1305139350757471055, language=CN, stringName=姚晓华, firstName=null, middleName=null, lastName=null, prefix=null, suffix=null, authorComment=null, nameInitials=null, affiliation=null, department=null, xref=^{2, 3}, address=^2．青海大学农林科学院，青海 西宁 810016
^3．青海省青稞遗传育种重点实验室/国家麦类改良中心青海青稞分中心/青藏高原种质资源研究与利用实验室，青海 西宁 810016, bio=null, bioImg=null, bioContent=null, aboutCorrespAuthor=null)}, companyList=[AuthorCompany(id=1305139347938898730, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, xref=2., ext=[AuthorCompanyExt(id=1305139347943093035, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, companyId=1305139347938898730, language=EN, country=null, province=null, city=null, postcode=null, companyName=null, departmentName=null, remark=^2.Qinghai University Academy of Agriculture and Forestry Sciences, Xining, Qinghai 810016, China), AuthorCompanyExt(id=1305139347951481644, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, companyId=1305139347938898730, language=CN, country=null, province=null, city=null, postcode=null, companyName=null, departmentName=null, remark=^2．青海大学农林科学院，青海 西宁 810016)]), AuthorCompany(id=1305139348018590509, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, xref=3., ext=[AuthorCompanyExt(id=1305139348144419630, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, companyId=1305139348018590509, language=EN, country=null, province=null, city=null, postcode=null, companyName=null, departmentName=null, remark=^3.Qinghai Provincial Key Laboratory of Qingke Genetic Breeding/Qinghai Barley Subcenter of National Wheat Improvement Center/Laboratory of Germplasm Resources Research and Utilization on the Qinghai-Tibet Plateau, Xining, Qinghai 810016, China), AuthorCompanyExt(id=1305139348157002543, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, companyId=1305139348018590509, language=CN, country=null, province=null, city=null, postcode=null, companyName=null, departmentName=null, remark=^3．青海省青稞遗传育种重点实验室/国家麦类改良中心青海青稞分中心/青藏高原种质资源研究与利用实验室，青海 西宁 810016)])])], keywords=[Keyword(id=1305139351021712212, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, language=EN, orderNo=1, keyword=Hulless barley), Keyword(id=1305139351080432469, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, language=EN, orderNo=2, keyword=Hinb-1 gene), Keyword(id=1305139351147541334, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, language=EN, orderNo=3, keyword=Grain hardness), Keyword(id=1305139351210455895, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, language=EN, orderNo=4, keyword=Cloning), Keyword(id=1305139351264981848, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, language=EN, orderNo=5, keyword=Expression analysis), Keyword(id=1305139351327896409, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, language=CN, orderNo=1, keyword=青稞), Keyword(id=1305139351395005274, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, language=CN, orderNo=2, keyword=Hinb-1基因), Keyword(id=1305139351457919835, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, language=CN, orderNo=3, keyword=籽粒硬度), Keyword(id=1305139351525028700, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, language=CN, orderNo=4, keyword=克隆), Keyword(id=1305139351579554653, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, language=CN, orderNo=5, keyword=表达分析)], refs=null, funds=[Fund(id=1305139351663440734, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, awardId=NKX202410, language=CN, fundingSource=青海大学农牧学院农科拔尖创新人才科研训练项目(NKX202410), fundOrder=null, country=null), Fund(id=1305139351734743903, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, awardId=null, language=CN, fundingSource=青海省青稞育种联合攻关项目, fundOrder=null, country=null), Fund(id=1305139351810241376, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, awardId=CARS-05, language=CN, fundingSource=国家大麦（青稞）产业技术体系专项(CARS-05), fundOrder=null, country=null)], companyList=[AuthorCompany(id=1305139347834041127, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, xref=1., ext=[AuthorCompanyExt(id=1305139347838235432, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, companyId=1305139347834041127, language=EN, country=null, province=null, city=null, postcode=null, companyName=null, departmentName=null, remark=^1.College of Agriculture and Animal Husbandry of Qinghai University, Xining, Qinghai 810016, China), AuthorCompanyExt(id=1305139347846624041, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, companyId=1305139347834041127, language=CN, country=null, province=null, city=null, postcode=null, companyName=null, departmentName=null, remark=^1．青海大学农牧学院，青海 西宁 810016)]), AuthorCompany(id=1305139347938898730, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, xref=2., ext=[AuthorCompanyExt(id=1305139347943093035, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, companyId=1305139347938898730, language=EN, country=null, province=null, city=null, postcode=null, companyName=null, departmentName=null, remark=^2.Qinghai University Academy of Agriculture and Forestry Sciences, Xining, Qinghai 810016, China), AuthorCompanyExt(id=1305139347951481644, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, companyId=1305139347938898730, language=CN, country=null, province=null, city=null, postcode=null, companyName=null, departmentName=null, remark=^2．青海大学农林科学院，青海 西宁 810016)]), AuthorCompany(id=1305139348018590509, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, xref=3., ext=[AuthorCompanyExt(id=1305139348144419630, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, companyId=1305139348018590509, language=EN, country=null, province=null, city=null, postcode=null, companyName=null, departmentName=null, remark=^3.Qinghai Provincial Key Laboratory of Qingke Genetic Breeding/Qinghai Barley Subcenter of National Wheat Improvement Center/Laboratory of Germplasm Resources Research and Utilization on the Qinghai-Tibet Plateau, Xining, Qinghai 810016, China), AuthorCompanyExt(id=1305139348157002543, tenantId=1146029695717560320, journalId=1303687221787852879, articleId=1305139346928071460, companyId=1305139348018590509, language=CN, country=null, province=null, city=null, postcode=null, companyName=null, departmentName=null, remark=^3．青海省青稞遗传育种重点实验室/国家麦类改良中心青海青稞分中心/青藏高原种质资源研究与利用实验室，青海 西宁 810016)])], figs=null, attaches=null, journal=Journal(id=1303686527903170636, delFlag=0, nameCn=麦类作物学报, nameEn=Journal of Triticeae Crops, nameHistory1=null, nameHistory2=null, issn=1009-1041, eissn=null, cn=61-1359/S, coden=null, periodic=0, language=CN, oaType=null, ccby=null, superviseOffice=null, ownerOffice=null, pubOffice=null, editorOffice=null, officeType=null, aims=null, clcCode=null, officeProv=null, officeCity=null, officeAddr=null, officeZip=null, officeEmail=null, officePhone=null, editDirector=null, officeDirector=null, officeDirectorPhone=null, officeStaffNum=null, officeEmpNum=null, coverPicUrl=/W3UvXD2ZVKDoQEt9zQGpg==, journalPrice=null, startedYear=null, abbrevIsoEn=Journal of Triticeae Crops, journalRemark=null, publicationField=null, createdTime=1788752689578, updatedTime=1788949384784, createdBy=18614031015, updatedBy=13041195026, firstLetterCn=M, firstLetterEn=M, subjectCode=Agriculture, subjectName=null, subjectCodeEn=Agriculture, subjectNameEn=null, picCn=/W3UvXD2ZVKDoQEt9zQGpg==, picEn=nyLiHfziMBV1Rm3HxncvkA==, jcr=null, cjcr=null, exts=[JournalExt(id=1304511528946921914, language=CN, name=麦类作物学报, nameHistory1=null, nameHistory2=null, managedBy=, sponsoredBy=, publishedBy=, editorOffice=, officeProv=null, officeCity=null, officeAddr=, officeZip=, editDirector=, officeDirector=null, officePhone=null, coverPicUrl=null, journalRemark=, submitArticleUrl=null, websiteUrl=, createdTime=1788949385153, updatedTime=1788949385153, createdBy=13041195026, updatedBy=13041195026, submissionGuidelinesUrl=, submissionAuthorUrl=http://mlzwxb.alljournals.ac.cn/mlzwxb/ch/author/login.aspx, submissionEditorUrl=http://mlzwxb.alljournals.ac.cn/mlzwxb/ch/login.aspx, submissionReviewUrl=http://mlzwxb.alljournals.ac.cn/mlzwxb/ch/auditor/login.aspx, submissionCeEditorUrl=, submissionAeEditorUrl=, option={"copyright":""}), JournalExt(id=1304511529018225083, language=EN, name=Journal of Triticeae Crops, nameHistory1=null, nameHistory2=null, managedBy=, sponsoredBy=, publishedBy=, editorOffice=, officeProv=null, officeCity=null, officeAddr=, officeZip=, editDirector=, officeDirector=null, officePhone=null, coverPicUrl=null, journalRemark=, submitArticleUrl=null, websiteUrl=, createdTime=1788949385170, updatedTime=1788949385170, createdBy=13041195026, updatedBy=13041195026, submissionGuidelinesUrl=, submissionAuthorUrl=http://mlzwxb.alljournals.ac.cn/mlzwxb/ch/author/login.aspx, submissionEditorUrl=http://mlzwxb.alljournals.ac.cn/mlzwxb/ch/login.aspx, submissionReviewUrl=http://mlzwxb.alljournals.ac.cn/mlzwxb/ch/auditor/login.aspx, submissionCeEditorUrl=, submissionAeEditorUrl=, option={"copyright":""})], databaseList=null, tenantJournalId=1303687221787852879, websiteList=[Website(id=1303713324831633955, webName=null, webTitle=null, webDomain=null, webCopyrigh=null, webIpcNo=null, seoTitle=null, seoKeywords=null, seoDescription=null, tenantJournalId=null, journalId=1303687221787852879, journalNameCn=null, journalNameEn=null, grayFlag=null, tenantId=1146029695717560320, platformId=null, journalGroupId=null, journalGroupNameCn=null, journalGroupNameEn=null, type=1, domain=https://castjournals.cast.org.cn/joweb/mlzwxb/CN, language=CN, createTime=1788759078462, createBy=18614031015, updateTime=1788759239556, updateBy=18614031015, name=麦类作物学报-中文, tplId=1146099689490845704, title=麦类作物学报, delFlag=0, indexPage=/home, props=[WebsiteProps(id=1303714116338741894, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1303713324831633955, code=articleTextType, value=kx, createTime=1788759267172, updateTime=1788759267172, creator=18614031015, updator=18614031015), WebsiteProps(id=1303714116246467203, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1303713324831633955, code=banner, value=null, createTime=1788759267150, updateTime=1788759267150, creator=18614031015, updator=18614031015), WebsiteProps(id=1303714116401656457, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1303713324831633955, code=grayFlag, value=0, createTime=1788759267187, updateTime=1788759267187, creator=18614031015, updator=18614031015), WebsiteProps(id=1303714116221301378, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1303713324831633955, code=logo, value=https://castjournals.cast.org.cn/joweb/mlzwxb/CN/file/pic?fileId=Ipfnv1HZql/CGnZchnsUyg==, createTime=1788759267144, updateTime=1788759267144, creator=18614031015, updator=18614031015), WebsiteProps(id=1303714116460376715, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1303713324831633955, code=minRunFlag, value=0, createTime=1788759267201, updateTime=1788759267201, creator=18614031015, updator=18614031015), WebsiteProps(id=1303714116309381765, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1303713324831633955, code=picServerUrl, value=https://castjournals.cast.org.cn/joweb/mlzwxb/CN/file/pic, createTime=1788759267165, updateTime=1788759267165, creator=18614031015, updator=18614031015), WebsiteProps(id=1303714116422627978, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1303713324831633955, code=silenceFlag, value=0, createTime=1788759267192, updateTime=1788759267192, creator=18614031015, updator=18614031015), WebsiteProps(id=1303714116280021636, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1303713324831633955, code=staticResourcePath, value=https://castjournals.cast.org.cn/joweb/cast_kjdb_cn_619/, createTime=1788759267158, updateTime=1788759267158, creator=18614031015, updator=18614031015), WebsiteProps(id=1303714116359713415, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1303713324831633955, code=themeColor, value=null, createTime=1788759267177, updateTime=1788759267177, creator=18614031015, updator=18614031015), WebsiteProps(id=1303714116380684936, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1303713324831633955, code=themeStyle, value=null, createTime=1788759267182, updateTime=1788759267182, creator=18614031015, updator=18614031015)]), Website(id=1303713324886159896, webName=null, webTitle=null, webDomain=null, webCopyrigh=null, webIpcNo=null, seoTitle=null, seoKeywords=null, seoDescription=null, tenantJournalId=null, journalId=1303687221787852879, journalNameCn=null, journalNameEn=null, grayFlag=null, tenantId=1146029695717560320, platformId=null, journalGroupId=null, journalGroupNameCn=null, journalGroupNameEn=null, type=1, domain=https://castjournals.cast.org.cn/joweb/mlzwxb/EN, language=EN, createTime=1788759078475, createBy=18614031015, updateTime=1788759235206, updateBy=18614031015, name=麦类作物学报-英文, tplId=1146101810881728533, title=Journal of Triticeae Crops, delFlag=0, indexPage=/home, props=[WebsiteProps(id=1303714087263826550, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1303713324886159896, code=articleTextType, value=kx, createTime=1788759260240, updateTime=1788759260240, creator=18614031015, updator=18614031015), WebsiteProps(id=1303714087234466419, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1303713324886159896, code=banner, value=null, createTime=1788759260233, updateTime=1788759260233, creator=18614031015, updator=18614031015), WebsiteProps(id=1303714087293186681, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1303713324886159896, code=grayFlag, value=0, createTime=1788759260247, updateTime=1788759260247, creator=18614031015, updator=18614031015), WebsiteProps(id=1303714087226077810, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1303713324886159896, code=logo, value=https://castjournals.cast.org.cn/joweb/mlzwxb/EN/file/pic?fileId=Ipfnv1HZql/CGnZchnsUyg==, createTime=1788759260231, updateTime=1788759260231, creator=18614031015, updator=18614031015), WebsiteProps(id=1303714087305769595, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1303713324886159896, code=minRunFlag, value=0, createTime=1788759260250, updateTime=1788759260250, creator=18614031015, updator=18614031015), WebsiteProps(id=1303714087255437941, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1303713324886159896, code=picServerUrl, value=https://castjournals.cast.org.cn/joweb/mlzwxb/EN/file/pic, createTime=1788759260238, updateTime=1788759260238, creator=18614031015, updator=18614031015), WebsiteProps(id=1303714087297380986, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1303713324886159896, code=silenceFlag, value=0, createTime=1788759260248, updateTime=1788759260248, creator=18614031015, updator=18614031015), WebsiteProps(id=1303714087247049332, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1303713324886159896, code=staticResourcePath, value=https://castjournals.cast.org.cn/joweb/cast_kjdb_en_623/, createTime=1788759260236, updateTime=1788759260236, creator=18614031015, updator=18614031015), WebsiteProps(id=1303714087272215159, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1303713324886159896, code=themeColor, value=null, createTime=1788759260242, updateTime=1788759260242, creator=18614031015, updator=18614031015), WebsiteProps(id=1303714087284798072, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1303713324886159896, code=themeStyle, value=null, createTime=1788759260245, updateTime=1788759260245, creator=18614031015, updator=18614031015)])], journalTitle=麦类作物学报, weixinUrl=null, journalUrl=http://mlzwxb.alljournals.ac.cn/, iacademicId=null, status=1, seqNo=null, journalTitleEn=Journal of Triticeae Crops, journalPhotoCn=/W3UvXD2ZVKDoQEt9zQGpg==, journalPhotoEn=nyLiHfziMBV1Rm3HxncvkA==, journalFirstLetter=M, journalRecommend=null, journalNew=null, journalCollection=null, jcrJf=null, cjcrJf=null, jcrJfStr=null, cjcrJfStr=null, submissionFirstDecision=null, sciSubjectClassification=null, casSubjectClassification=null, citeScore=null, totalCitationFrequency=null, icpCode=null, psCode=null, advertisingLicenseCode=null, copyrightInformation=null, country=null, option=, provinceCode=null, provinceName=null, collectFlag=false, interPubPlatform=, interPubPlatformUrl=null), detailUrlCn=https://castjournals.cast.org.cn/joweb/mlzwxb/CN/10.7606/j.issn.1009-1041.2026.03.04, detailUrlEn=https://castjournals.cast.org.cn/joweb/mlzwxb/EN/10.7606/j.issn.1009-1041.2026.03.04, pdfUrlCn=https://castjournals.cast.org.cn/joweb/mlzwxb/CN/PDF/10.7606/j.issn.1009-1041.2026.03.04, pdfUrlEn=https://castjournals.cast.org.cn/joweb/mlzwxb/EN/PDF/10.7606/j.issn.1009-1041.2026.03.04, aliStartDate=0, aliEndDate=0, collectionFlag=false, citedCount=null, citedUrl=null, previewStatus=0, delFlag=0, hasFullText=0, orderTime=1773504000000, fullTextJson=null, articleText=null, reference=null)