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Blackburn, Carol W. Greider and Jack W. Szostak for their discovery of how chromosomes are protected by telomeres and the enzyme telomerase. Telomere is a specific structure of chromosomal termini composed of a series of DNA repeats, to protect the stability and integrity of chromosome. Depending on an intrinsic RNA template, telomerase synthesizes telomere DNA, and maintains the length of telomere. The discovery of telomere and telomerase has solved a long-standing fundamental problem in biology: how can the end of chromosomes be maintained and spared from erosion during repeated cellular divisions. 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