菌物学报

中国贵州省木生大型担子菌编目

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赵恒 ¹^,² , 陈芊 ³ , 张欣 ¹ , 罗凯月 ¹ , 李彪 ¹ , 员瑗 ¹ , 吴芳 ¹ , 何双辉 ¹ , 曾广宇 ¹^,⁴ , 庄磊 ¹^,⁵ , 戴玉成 ¹^,^* , 邓春英 ⁶^,^*

菌物学报 | 研究论文 2026,45(4): 250358

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菌物学报 | 研究论文 2026, 45(4): 250358

中国贵州省木生大型担子菌编目

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赵恒¹^,², 陈芊³, 张欣¹, 罗凯月¹, 李彪¹, 员瑗¹, 吴芳¹, 何双辉¹, 曾广宇¹^,⁴, 庄磊¹^,⁵, 戴玉成¹^,^*, 邓春英⁶^,^*

作者信息

¹ 北京林业大学生态与自然保护学院，北京 100083

² 中国科学院沈阳应用生态研究所森林生态与保育重点实验室，辽宁沈阳 110016

³ 重庆交通大学建筑与城市规划学院，重庆 400074

⁴ 广西壮族自治区林业科学研究院，广西南宁 530002

⁵ 哈尔滨市农业科学院，黑龙江哈尔滨 150028

⁶ 贵州科学院贵州省生物研究所，贵州贵阳 550009

A checklist of lignicolous macro-basidiomycetes in Guizhou Province, southwest China

Heng ZHAO¹^,², Qian CHEN³, Xin ZHANG¹, Kaiyue LUO¹, Biao LI¹, Yuan YUAN¹, Fang WU¹, Shuanghui HE¹, Guangyu ZENG¹^,⁴, Lei ZHUANG¹^,⁵, Yucheng DAI¹^,^*, Chunying DENG⁶^,^*

Affiliations

¹ School of Ecology and Nature Conservation, Beijing Forestry University, Beijing 100083, China

² CAS Key Laboratory of Forest Ecology and Silviculture, Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang 110016, Liaoning, China

³ College of Architecture and Urban Planning, Chongqing Jiaotong University, Chongqing 400074, China

⁴ Guangxi Forestry Science Research Institute, Nanning 530002, Guangxi, China

⁵ Harbin Academy of Agricultural Sciences, Harbin 150028, Heilongjiang, China

⁶ Guizhou Institute of Biology, Guizhou Academy of Sciences, Guiyang 550009, Guizhou, China

出版时间: 2026-04-22 doi: 10.13346/j.mycosystema.250358

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摘要

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贵州省位于云贵高原东部，地形复杂、气候温暖湿润，具有丰富的真菌多样性潜力，但其木生大型担子菌资源长期缺乏系统梳理。本研究基于4个研究团队的数据，特别是近年在贵州省采集标本的深入研究，并结合发表的文献资料，对贵州省木生大型担子菌多样性进行了系统整理与统计。结果表明，贵州省木生大型担子菌共计738种，隶属于担子菌门3纲16目74科253属，其中担子菌纲为绝对优势类群，占总物种数的97.15%。在目级水平上，多孔菌目、蘑菇目和锈革孔菌目为优势目，共包含601种，占总物种数的81.44%。科级水平上，10个优势科共包含391种，占总物种数的52.98%；属级水平上，11个优势属共计202种，占总物种数的27.37%。此外，共有48种以贵州省作为模式或副模式产地。本研究在省域尺度系统揭示了贵州省木生大型担子菌的物种组成和多样性格局。

关键词

木生真菌 / 大型真菌 / 真菌区系 / 特种多样性

Abstract

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Guizhou Province, located in the eastern Yunnan-Guizhou Plateau, has complex topography and a warm, humid climate, providing high potential for fungal diversity. However, lignicolous macro-basidiomycetes in this region lacked systematic assessment for a long time. A comprehensive compilation of lignicolous macro-basidiomycetes in Guizhou was conducted based on specimen records of four investigation teams and recently published literature. A total of 738 species were recorded, belonging to 3 classes, 16 orders, 74 families, and 253 genera within Basidiomycota. The dominant class is Agaricomycetes, with the sum of species accounting for 97.15% of all known lignicolous macro-basidiomycetes species in Guizhou. The dominant orders are Polyporales, Agaricales, and Hymenochaetales, and the sum of species accounts for 81.44% (601 species) of all lignicolous macro-basidiomycetes in Guizhou. The ten dominant families include 391 species, accounting for 52.98% of total known species. The eleven dominant genera consist of 202 species, accounting for 27.37% of the total known species. Additionally, 48 species are described on the basis of holotype or paratype collected from Guizhou. This is the first province-level systematic overview of lignicolous macro-basidiomycetes in Guizhou Province.

Key words

wood-inhabiting fungi / macrofungi / funga / species diversity

引用本文

赵恒, 陈芊, 张欣, 罗凯月, 李彪, 员瑗, 吴芳, 何双辉, 曾广宇, 庄磊, 戴玉成, 邓春英. 中国贵州省木生大型担子菌编目. 菌物学报, 2026 , 45 (4) : 250358 - . DOI: 10.13346/j.mycosystema.250358

Heng ZHAO, Qian CHEN, Xin ZHANG, Kaiyue LUO, Biao LI, Yuan YUAN, Fang WU, Shuanghui HE, Guangyu ZENG, Lei ZHUANG, Yucheng DAI, Chunying DENG. A checklist of lignicolous macro-basidiomycetes in Guizhou Province, southwest China[J]. Mycosystema, 2026 , 45 (4) : 250358 - . DOI: 10.13346/j.mycosystema.250358

正文

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木生大型担子菌是指在森林生态系统中以森林生态系统为主要生境，以木材(包括活立木或枯木)主要成分作为主要营养来源的一类隶属于担子菌门的大型真菌。该类群能够在木质纤维素基质上生长、定殖、产生生物酶，从而引起木材腐朽，部分物种还可与活树形成寄生或共生关系，在森林生态系统的更新过程以及碳、氮等生物地球化学循环中发挥着不可替代的重要作用(司静等2011；Wang et al. 2013；戴玉成等 2021；Spirin et al. 2024；Zhao et al. 2024)。在生态功能上，木生大型担子菌的主要类群是木材腐朽真菌，根据其对木材细胞壁主要成分的降解能力不同，可主要被划分为白腐真菌(white rot)和褐腐真菌(brown rot) (Krah et al. 2018)。此外，部分木生大型担子菌是树木的重要病原菌，能够侵染活立木并引发心材腐朽、根腐病和干腐病等多种森林病害，对森林健康与木材资源造成显著影响(Yuan et al. 2021, 2023；Wu et al. 2022)。与此同时，木生大型担子菌还是重要的食用和药用真菌资源之一，具有悠久的利用历史，例如黑木耳Auricularia heimuer、杨树桑黄Sanghuangporus vaninii和灵芝Ganoderma lingzhi等，在食品安全、医药开发和生物产业中具有重要价值(Si et al. 2019；Wu et al. 2019；吴声华和戴玉成 2020)。

近年来，中国学者围绕木生大型真菌多样性开展了大量系统的资源调查与区系特征研究，研究尺度涵盖单一省份、山脉、自然保护区乃至全国范围，显著推进了中国木生真菌分类学与生物地理学研究(Dai 2012；董军红等2023；Dong et al. 2024；王科等2024；朱安红等2024；马玉硕等2025；Xu et al. 2025)。例如，袁海生等(2009)在黑龙江小兴安岭东部地区报道木生真菌124种，其中包括105种多孔菌和19种革菌；Ma et al. (2022)报道了海南木生真菌702种；边禄森等(2023)在太行山地区鉴定出隶属于2门4纲12目49科147属，共306种木生真菌；周林江等(2023)在山东省共记录木材腐朽真菌隶属于2门6纲15目54科131属和227种。这些研究为认识中国木生真菌的物种多样性和区系格局奠定了重要基础。

贵州省位于中国西南地区，地处长江与珠江上游水系交汇地带，属于云贵高原东部的典型内陆山地省份。该区域地形复杂、气候温暖湿润，属亚热带季风气候类型，孕育了高度异质的森林生态系统。其中，梵净山等区域被认为是中国西南部生物多样性热点之一，是开展森林生态学、真菌多样性及生态保护研究的重要区域。贵州省在大型真菌多样性研究方面已取得一定进展。例如，《中国真菌总汇》记录的6 737个物种中分布在贵州省的担子菌门真菌约180种(戴芳澜1979)；姚刘斌等(2018)在正安县记录菌根真菌30种，隶属于15科19属；王晶等(2024)在武陵山区西部优先保护区报道大型真菌隶属于2门13纲26目91科304属和1 148种；杨云深等(2025)在桴焉自然保护区调查发现大型真菌隶属于5纲26目70科152属和392种。这些研究结果表明，贵州省具有极为丰富的大型真菌物种多样性。

然而，迄今为止，贵州省木生大型担子菌多样性的本底资源调查仍显不足。现有关于该类群的研究多以个别区域或特定类群为对象，相关物种记录零散分布于不同文献之中，尚缺乏全省尺度的系统整理与综合总结(姚刘斌等 2018；邓春英等2022, 2025；Wu et al. 2022；王晶等 2024；Liu et al. 2025a, 2025b；Qin et al. 2025；杨云深等 2025；Zhao et al. 2025)。这一现状在一定程度上制约了贵州省木生真菌资源的科学认识及其在林业、食药用菌和相关生物产业中的合理开发与利用。因此，系统厘清贵州省木生大型担子菌的物种多样性具有重要意义。

本研究基于我国4个研究团队的协同合作下，系统整合北京林业大学微生物研究所标本馆(Herbarium of the Institute of Microbiology, Beijing Forestry University, BJFC)和中国科学院沈阳应用生态研究所东北生物标本馆(Herbarium of Northeast China, Institute of Applied Ecology, Chinese Academy of Sciences, IFP)馆藏数据，鉴定了作者近年在贵州省采集的标本，结合近年来发表的相关文献记录，对贵州省木生大型担子菌多样性进行了全面梳理与总结。研究共确认贵州省木生大型担子菌738种，其中以贵州省作为模式产地的物种共有48种，为深入认识该区域木生大型担子菌多样性和资源潜力奠定了坚实基础。

1 数据的基本信息

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本研究数据的名称为中国贵州省木生大型担子菌编目。数据的作者是赵恒，陈芊，张欣，罗凯月，李彪，员瑗，吴芳，何双辉，曾广宇，庄磊，戴玉成，邓春英。数据产生的时间是2025年12月10日。数据量是738种贵州省木生大型担子菌物种名录。数据格式为word形式，数据提交至科学数据银行(Science Data Bank)期刊社区(https://scidb.cn/jwxb)，CSTR编号：31253.11.sciencedb.36968，DOI编号：10.57760/ sciencedb.36968。

2 数据采集与处理方法

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2.1 数据研究区域概括

研究区域覆盖贵州省全境。贵州省位于中国西南部、云贵高原东部，行政区总面积约为17.6万km²，地理坐标范围为103°36′-109°35′E、24°37′-29°13′N。省内地势起伏显著，海拔高度介于147.8-2 900 m之间，地形以高原、山地和丘陵为主。贵州省东邻湖南省，南接广西壮族自治区，西与云南省相连，北与四川省及重庆市相毗，共同构成中国西南地区重要的生态单元。

2.2 研究材料和数据来源

研究材料来源于北京林业大学、中国科学院沈阳应用生态研究所、重庆交通大学和贵州科学院贵州省生物研究所4个研究团队从2000年开始对贵州省的多次大型真菌资源野外采集，主要的调查点包括：安顺市西秀区甘堡村场、安顺市西秀区老落坡林场、安顺市黄果树瀑布、八舟河国家湿地公园、百花湖森林公园、毕节市白马山林场、毕节市百里杜鹃风景区、毕节市拱拢坪国家森林公园、毕节市赫章县赫章国家森林公园、毕节市赫章县水塘林场、毕节市威宁县小海镇、毕节市织金县织金洞门口、赤水市桫椤自然保护区、贵阳阿哈湖国家湿地公园、贵阳喀斯特公园、贵阳市扁坡林场、贵阳市贵州师范大学校内、贵阳市贵州植物园、贵阳市河滨公园、贵阳市黔灵山公园、贵阳市森林公园、贵阳市桃源河景区、贵阳市西望山、贵州省农业科学院、贵州省植物园、江口县亚木沟风景区、雷公山国家森林公园、黎平国家森林公园、荔波县观音坡、荔波县茂兰国家自然保护区、荔波县小七孔景区、六盘水市水城县芍米乡玉舍国家森林公园、六盘水市玉舍国家森林公园、龙架山国家森林公园、鹿冲关森林公园、麻江县坝茫乡老蛇冲自然保护区、茂兰喀斯特森林、梅兰山公园、南明区森林公园、盘龙山森林公园、骑龙山人文生态纪念园、黔南布依族苗族自治州斗篷山国家自然保护区、黔西南州普晴林场、清镇市九龙山、榕江县沙坪沟护林站、施秉县云台山、顺海公园、绥阳县宽阔水自然保护区、太平山森林公园、铜仁市江口县梵净山保护区、望谟县望谟自然保护区、威宁彝族回族苗族自治县草海国家自然保护区、乌江白果坨国家湿地公园、习水县习水国家自然保护区、仙女潭保护区、兴义市丰都林场、兴义市马岭河、印江县梵净山风景区、长坡岭国家森林公园等地区共计收集标本3 000余份。此外，本研究同时汇总最新文献报道(Zhang & Yao 2019；戴玉成等2021；Li et al. 2021, 2022a, 2022b；Li et al. 2025；邓春英等2022, 2025；Deng et al. 2024, 2025；Zhu et al. 2024；Liu et al. 2025a, 2025b；Qin et al. 2025；Wang et al. 2025；Xu et al. 2025；杨云深等2025；Zhao et al. 2025)。

2.3 统计分析

本研究中基于形态学和分子生物学对采集的标本进行鉴定，同时使用Index Fungorum数据库(http://www.indexfungorum.org/Names/Names.asp)和MycoBank数据库(https://www.mycobank. org/Simple%20names%20search)对所有物种名称进行校对整理，排除错误物种名称和同物异名，得到中国贵州省木生大型担子菌物种名录。此外，基于该物种名录，并参考已有的研究方法(王妍等 2021；孙渤洋等 2023)，统计分析优势目(物种数量大于等于50种)、优势科(物种数量大于等于20种)和优势属(物种数量大于等于10种)。

2.4 中国贵州省木生大型担子菌目录

本研究共收集并整理贵州省木生大型担子菌738种，其中48种以贵州省作为其模式产地(holotype)或副模式产地(paratype)，并以“*”号加以标识，部分物种生境新鲜子实体见图1。物种名录按照属的拉丁学名首字母顺序进行排列，具体如下。

1. Abortiporus biennis (Bull.) Singer

2. Abundisporus pubertatis (Lloyd) Parmasto

3. Agrocybe parasitica G. Stev.

4. Alboefibula gracilis C.C. Chen & Sheng H. Wu

5. Aleurocystidiellum tsugae (Yasuda) S.H. He & Y.C. Dai

6. Aleurodiscus botryosus Burt

7. Aleurodiscus cerussatus (Bres.) Höhn. & Litsch.

8. Aleurodiscus rimulosus Núñez & Ryvarden

9. Aleurodiscus subroseus S.H. He & Y.C. Dai

10. Aleurodiscus wakefieldiae Boidin & Beller

11. Alloexidiopsis calcea (Pers.) L.W. Zhou & S.L. Liu

12. Alloexidiopsis grandinea J.H. Dong & C.L. Zhao

13. Alloexidiopsis sinensis J.H. Dong & C.L. Zhao

14. Allophlebia formosana (Sheng H. Wu) S.H. He et al.

15. Amaropostia stiptica (Pers.) B.K. Cui et al.

16. Antella americana (Ryvarden & Gilb.) Ryvarden

17. Antrodia albida (Fr.) Donk

18. Antrodia macrospora Bernicchia & De Dominicis

19. Antrodia sinuosa (Fr.) P. Karst.

20. Antrodiella lactea H.S. Yuan

21. Antrodiella onychoides (Egeland) Niemelä

22. Antrodiella parasitica Vampola

23. Antrodiella semisupina (Berk. & M.A. Curtis) Ryvarden

24. Armillaria amygdalispora H.C. Wang et al. *

25. Armillaria cepistipes Velen.

26. Armillaria gallica Marxm. & Romagn.

27. Armillaria mellea (Vahl) P. Kumm.

28. Asterostroma cervicolor (Berk. & M.A. Curtis) Massee

29. Asterostroma muscicola (Berk. & M.A. Curtis) Massee

30. Asterostroma rhizomorpharum H.M. Zhou & C.L. Zhao *

31. Athelia acrospora Jülich

32. Aurantiporus transformatus (Núñez & Ryvarden) Spirin & Zmitr.

33. Auricularia cornea Ehrenb.

34. Auricularia delicata (Mont. ex Fr.) Henn.

35. Auricularia fibrillifera Kobayasi

36. Auricularia fuscosuccinea (Mont.) Henn.

37. Auricularia heimuer F. Wu et al.

38. Auricularia nigricans (Sw.) Birkebak et al.

39. Auricularia sinodelicata Y.C. Dai & F. Wu

40. Auricularia villosula Malysheva

41. Auriporia aurulenta A. David et al.

42. Auriscalpium vulgare Gray

43. Bjerkandera adusta (Willd.) P. Karst.

44. Bjerkandera fumosa (Pers.) P. Karst.

45. Boidinia donkii (S.S. Rattan) Sheng H. Wu & P.K. Buchanan

46. Boreostereum vibrans (Berk. & M.A. Curtis) Davydkina & Bondartseva

47. Botryobasidium coniferarum S.L. Liu & L.W. Zhou *

48. Botryobasidium indicum (P.N. Singh & S.K. Singh) R. Kirschner & G. Langer

49. Butyrea japonica (Núñez & Ryvarden) Miettinen & Ryvarden

50. Byssomerulius corium (Pers.) Parmasto

51. Cabalodontia albofibrillosa (Hjortstam & Ryvarden) Westphalen

52. Calocera cornea (Batsch) Fr.

53. Calocera viscosa (Pers.) Bory

54. Callistosporium luteo-olivaceum (Berk. & M.A. Curtis) Singer

55. Campanella buettneri Henn.

56. Campanella junghuhnii (Mont.) Singer

57. Candolleomyces candolleanus (Fr.) D. Wächt. & A. Melzer

58. Candolleomyces parvipileus Javaid & Naseer

59. Cerioporus scutellatus (Schwein.) Zmitr.

60. Cerioporus subtropicus (B.K. Cui et al.) Zmitr.

61. Cerioporus varius (Pers.) Zmitr. & Kovalenko

62. Ceriporia aurantiocarnescens (Henn.) M. Pieri & B. Rivoire

63. Ceriporia excelsa Parmasto

64. Ceriporia mellita (Bourdot & Galzin) Bondartsev & Singer

65. Ceriporia occidentalis Spirin & Vlasák

66. Ceriporia subbadia (Murrill) Y.C. Dai et al.

67. Ceriporia viridans (Berk. & Broome) Donk

68. Ceriporiopsis mucida (Pers.) Gilb. & Ryvarden

69. Ceriporiopsis subvermispora (Pilát) Gilb. & Ryvarden

70. Cerrena albocinnamomea (Y.C. Dai & Niemelä) H.S. Yuan

71. Cerrena unicolor (Bull.) Murrill

72. Cerrena zonata (Berk.) H.S. Yuan

73. Chaetocalathus craterellus (Durieu & Lév.) Singer

74. Chaetocalathus galeatus (Berk. & M.A. Curtis) Singer

75. Cheimonophyllum haedinum (Berk. & M.A. Curtis) Valade & P.-A. Moreau

76. Chondrostereum purpureum (Pers.) Pouzar

77. Cinereomyces lindbladii (Berk.) Jülich

78. Clitocybe nebularis (Batsch) P. Kumm.

79. Collybia nivea (Mont.) Dennis

80. Collybiopsis biformis (Peck) R.H. Petersen

81. Collybiopsis confluens (Pers.) R.H. Petersen

82. Collybiopsis dichroa (Berk. & M.A. Curtis) R.H. Petersen

83. Collybiopsis foliiphila (A.K. Dutta et al.) R.H. Petersen

84. Collybiopsis luxurians (Peck) R.H. Petersen

85. Collybiopsis menehune (Desjardin et al.) R.H. Petersen

86. Collybiopsis nonnulla (Corner) R.H. Petersen

87. Collybiopsis peronata (Bolton) R.H. Petersen

88. Collybiopsis polygramma (Mont.) R.H. Petersen

89. Collybiopsis ramealis (Bull.) Millsp.

90. Collybiopsis stenophylla (Mont.) R.H. Petersen

91. Collybiopsis subnuda (Ellis ex Peck) R.H. Petersen

92. Coltricia baoshanensis (Y.C. Dai & B.K. Cui) Y.C. Dai & F. Wu

93. Coltricia cinnamomea (Jacq.) Murrill

94. Coltricia crassa Y.C. Dai

95. Coltricia minima L.S. Bian & Y.C. Dai

96. Coltricia perennis (L.) Murrill

97. Coltricia pusilla Imazeki & Kobayasi

98. Coltricia spina Y.C. Dai

99. Coltricia subperennis (Z.S. Bi & G.Y. Zheng) G.Y. Zheng & Z.S. Bi

100. Coltricia subpicta (Lloyd) Imazeki & Kobayasi

101. Coltricia strigosipes Corner

102. Coltricia subpicta (Lloyd) Imazeki & Kobayasi

103. Coltricia subverrucata L.S. Bian & Y.C. Dai *

104. Coltricia tsugicola Y.C. Dai & B.K. Cui

105. Coltricia verrucata Aime et al.

106. Coltricia weii Y.C. Dai

107. Coprinopsis rhombisporoides Voto & Deschuyteneer

108. Cotylidia aurantiaca (Pat.) A.L. Welden

109. Cotylidia diaphana (Cooke) Lentz

110. Crepidotus applanatus (Pers.) P. Kumm.

111. Crepidotus crocophyllus (Berk.) Sacc.

112. Crepidotus dentatus T. Bau & Y.P. Ge

113. Crepidotus mollis (Schaeff.) Staude

114. Crepidotus parietalis E. Horak

115. Crepidotus stenocystis Pouzar

116. Crepidotus subverrucisporus Pilát

117. Crepidotus sulphurinus Imazeki & Toki

118. Crepidotus variabilis (Pers.) P. Kumm.

119. Crucibulum laeve (Huds.) Kambly

120. Crustomyces isabellinus (Berk. & Broome) Yue Li et al.

121. Cryptoporus sinensis Sheng H. Wu & M. Zang

122. Cubamyces lactineus (Berk.) Lücking

123. Cyanosporus alni (Niemelä & Vampola) B.K. Cui et al.

124. Cyanosporus caesius (Schrad.) McGinty

125. Cyanosporus coeruleivirens (Corner) B.K. Cui et al.

126. Cyanosporus fusiformis B.K. Cui et al. *

127. Cyanosporus glaucus (Spirin & Miettinen) B.K. Cui & Shun Liu

128. Cyanosporus simulans (P. Karst.) B.K. Cui & Shun Liu

129. Cyanosporus subhirsutus B.K. Cui et al. *

130. Cyanosporus tabuliformis Y.C. Dai et al.

131. Cyanosporus tricolor B.K. Cui et al.

132. Cyanotrama rimosa (Murrill) Ghob.-Nejh. & Y.C. Dai

133. Cyathus striatus Willd.

134. Cylindrobasidium laeve (Pers.) Chamuris

135. Dacrymyces capitatus Schwein.

136. Dacrymyces chrysospermus Berk. & M.A. Curtis

137. Dacrymyces spathularia (Schwein.) Alvarenga

138. Dacrymyces stillatus Nees

139. Dacryobolus karstenii (Bres.) Oberw. ex Parmasto

140. Dacryobolus montanus X.Z. Wan & H.S. Yuan

141. Dacryobolus sudans (Alb. & Schwein.) Fr.

142. Dacryopinax sphenocarpa Shirouzu & Tokum.

143. Daedalea xantha (Fr.) A. Roy & A.B. De

144. Daedaleopsis confragosa (Bolton) J. Schröt.

145. Daedaleopsis nipponica Imazeki

146. Daedaleopsis tricolor (Bull.) Bondartsev & Singer

147. Datronia perstrata (Corner) T. Hatt. & Sotome

148. Dentipellis longiuscula L.L. Shen & Min Wang

149. Desarmillaria tabescens (Scop.) R.A. Koch & Aime

150. Diplomitoporus flavescens (Bres.) Domański

151. Ditiola peziziformis (Lév.) D.A. Reid

152. Duportella tristicula (Berk. & Broome) Reinking

153. Earliella scabrosa (Pers.) Gilb. & Ryvarden

154. Echinochaete russiceps (Berk. & Broome) D.A. Reid

155. Eichleriella discolor (Berk. & Broome) S.H. He & Nakasone

156. Eichleriella shearii (Burt) Spirin & Malysheva

157. Eichleriella sinensis (Teng) S.H. He & Nakasone

158. Eichleriella tenuicula (Lév.) Spirin & Malysheva

159. Eichleriella xinpingensis C.L. Zhao

160. Elmerina caryae (Schwein.) D.A. Reid

161. Exidia glandulosa (Bull.) Fr.

162. Exidia punctata Lu Wang & C.L. Zhao

163. Exidia uvapassa Lloyd

164. Fabisporus sanguineus (L.) Zmitr.

165. Favolaschia nipponica Kobayasi

166. Favolaschia sprucei (Berk.) Singer

167. Favolaschia tonkinensis (Pat.) Kuntze

168. Favolus grammocephalus (Berk.) Imazeki

169. Favolus tenuiculus P. Beauv.

170. Fibroporia radiculosa (Peck) Parmasto

171. Fistulina hepatica (Schaeff.) With.

172. Flammulina filiformis (Z.W. Ge et al.) P.M. Wang et al.

173. Flavodon flavus (Klotzsch) Ryvarden

174. Flammula alnicola (Fr.) P. Kumm.

175. Fomitiporella sinica Y.C. Dai et al.

176. Fomitiporia punctata (P. Karst.) Murrill

177. Fomitiporia torreyae Y.C. Dai & B.K. Cui

178. Fomitopsis betulina (Bull.) B.K. Cui et al.

179. Fomitopsis malicola (Berk. & M.A. Curtis) Spirin

180. Fomitopsis modesta (Kuntze ex Fr.) Vlasák & Spirin

181. Fomitopsis nigra (Berk.) Spirin & Miettinen

182. Fomitopsis nivosa (Berk.) Gilb. & Ryvarden

183. Fomitopsis palustris (Berk. & M.A. Curtis) Gilb. & Ryvarden

184. Fomitopsis pinicola (Sw.) P. Karst.

185. Fomitopsis pulvina (Pers.) V. Spirin & J. Vlasák

186. Fomitopsis pulvinascens (Pilát) T. Niemelä & O. Miettinen

187. Foraminispora concentrica (J. Song et al.) Y.F. Sun & B.K. Cui

188. Fragiliporia fragilis Y.C. Dai et al.

189. Fulvifomes inermis (Ellis & Everh.) Y.C. Dai

190. Fulvifomes merrillii (Murrill) Baltazar & Gibertoni

191. Fulvoderma australe L.W. Zhou & Y.C. Dai

192. Fulvoderma scaurum (Lloyd) L.W. Zhou & Y.C. Dai

193. Funalia hispida (Bagl.) M.M. Chen

194. Funalia leonina (Klotzsch) Pat.

195. Funalia sanguinaria (Klotzsch) Zmitr. & Malysheva

196. Fuscoporia chinensis Q. Chen et al.

197. Fuscoporia contigua (Pers.) G. Cunn.

198. Fuscoporia ferrea (Pers.) G. Cunn.

199. Fuscoporia gilva (Schwein.) T. Wagner & M. Fisch.

200. Fuscoporia karsteniana Q. Chen et al.

201. Fuscoporia monticola Y.C. Dai et al.

202. Fuscoporia rhabarbarina (Berk.) Groposo et al.

203. Fuscoporia subferrea Q. Chen bis & Yuan Yuan

204. Fuscopostia fragilis (Fr.) B.K. Cui et al.

205. Fuscopostia leucomallella (Murrill) B.K. Cui et al.

206. Ganoderma applanatum (Pers.) Pat.

207. Ganoderma australe (Fr.) Pat.

208. Ganoderma duropora Lloyd

209. Ganoderma flexipes Pat.

210. Ganoderma gibbosum (Blume & T. Nees) Pat.

211. Ganoderma lingzhi Sheng H. Wu et al.

212. Ganoderma lucidum (Curtis) P. Karst.

213. Ganoderma orbiforme (Fr.) Ryvarden

214. Ganoderma ovisporum H.D. Yang & T.C. Wen *

215. Ganoderma sanduense Hapuar. et al. *

216. Ganoderma sichuanense J.D. Zhao & X.Q. Zhang

217. Ganoderma sinense J.D. Zhao et al.

218. Ganoderma tsugae Murrill

219. Gerronema kuruvense K.P.D. Latha & Manim.

220. Gerronema strombodes (Berk. & Mont.) Singer

221. Gerronema subclavatum (Peck) Singer ex Redhead

222. Gloeocystidiellum porosum (Berk. & M.A. Curtis) Donk

223. Gloeocystidiopsis heimii (Boidin) Jülich

224. Gloeophyllum sepiarium (Wulfen) P. Karst.

225. Gloeophyllum striatum (Fr.) Murrill

226. Gloeosoma mirabile (Berk. & M.A. Curtis) Rajchenb. et al.

227. Grifola huishuhua X.J. Xie et al.

228. Guepinia helvelloides (DC.) Fr.

229. Guepiniopsis buccina (Pers.) L.L. Kenn.

230. Gymnopilus aurantiacus Hesler

231. Gymnopilus dryophilus Murrill

232. Gymnopilus junonius (Fr.) P.D. Orton

233. Gymnopilus liquiritiae (Pers.) P. Karst.

234. Gymnopilus penetrans (Fr.) Murrill

235. Gymnopilus terrestris Hesler

236. Gymnopus androsaceus (L.) Della Magg. & Trassin.

237. Gymnopus efibulatus J.P. Li et al. *

238. Gymnopus erythropus (Pers.) Antonín et al.

239. Gymnopus fuscopurpureus (Pers.) Antonín et al.

240. Gymnopus omphalinoides J.P. Li *

241. Gymnopus pallipes J.P. Li & Chun Y. Deng *

242. Gymnopus pseudoandrosaceus Chun Y. Deng et al. *

243. Gymnopus strigosipes J.P. Li et al. *

244. Haddowia longipes (Lév.) Steyaert

245. Hapalopilus rutilans (Pers.) Murrill

246. Haploporus alabamae (Berk. & Cooke) Y.C. Dai & Niemelä

247. Haploporus latisporus Juan Li & Y.C. Dai

248. Haploporus papyraceus (Cooke) Y.C. Dai & Niemelä

249. Haploporus subtrameteus (Pilát) Y.C. Dai & Niemelä

250. Hemimycena oregonensis (A.H. Sm.) Singer

251. Hericium erinaceus (Bull.) Pers.

252. Heterobasidion insulare (Murrill) Ryvarden

253. Heterochaete falcato-sporifera Y.B. Peng & X.W. Hu

254. Heterochaete roseola Pat.

255. Heterochaete sinensis Teng

256. Heterocorticium bambusicola S.H. He et al.

257. Heteroradulum labyrinthinum (H.S. Yuan & Decock) L.W. Zhou

258. Heteroradulum maolanense S.H. He et al. *

259. Hirschioporus abietinus (Pers. ex J.F. Gmel.) Donk

260. Hirschioporus fuscoviolaceus (Ehrenb.) Donk

261. Hohenbuehelia grisea (Peck) Singer

262. Hohenbuehelia petaloides (Bull.) Schulzer

263. Hydnochaete japonica Lloyd

264. Hydnophlebia acanthocystis (Gilb. & Nakasone) C.L. Zhao

265. Hydnoporia conifera S.H. He et al. *

266. Hydnoporia corrugata (Fr.) K.H. Larss. & Spirin

267. Hydnoporia subrigidula (S.H. He & Hai J. Li) Miettinen & K.H. Larss.

268. Hydnoporia tabacinoides (Yasuda) Miettinen & K.H. Larss.

269. Hydnoporia yasudae (Imazeki) Spirin & Miettinen

270. Hymenochaete anomala Burt

271. Hymenochaete austrosinensis S.H. He et al. *

272. Hymenochaete cinnamomea (Pers.) Bres.

273. Hymenochaete coffeana J.C. Léger & Lanq.

274. Hymenochaete cruenta (Pers.) Donk

275. Hymenochaete flava S.H. He et al. *

276. Hymenochaete floridea Berk. & Broome

277. Hymenochaete hydnoides T. Wagner & M. Fisch.

278. Hymenochaete innexa G. Cunn.

279. Hymenochaete luteomarginata S.H. He et al. *

280. Hymenochaete microcycla (Zipp. ex Lév.) Spirin & Miettinen

281. Hymenochaete minor S.H. He & Y.C. Dai

282. Hymenochaete minuscula G. Cunn.

283. Hymenochaete moniliformis S.H. He et al.

284. Hymenochaete mougeotii (Fr.) Cooke

285. Hymenochaete muroiana I. Hino & Katum.

286. Hymenochaete nanospora J.C. Léger

287. Hymenochaete ochromarginata P.H.B. Talbot

288. Hymenochaete parmastoi S.H. He & Hai J. Li

289. Hymenochaete ramicola S.H. He et al. *

290. Hymenochaete rheicolor (Mont.) Lév.

291. Hymenochaete rubiginosa (Dicks.) Lév.

292. Hymenochaete separabilis J.C. Léger

293. Hymenochaete setulohypha S.H. He et al. *

294. Hymenochaete sphaeriicola Lloyd

295. Hymenochaete subfissurata S.H. He et al. *

296. Hymenochaete xerantica (Berk.) S.H. He & Y.C. Dai

297. Hymenochaete villosa (Lév.) Bres.

298. Hyphoderma ayresii (Berk. ex Cooke) Boidin & Gilles

299. Hyphoderma crystallinum C.L. Zhao & Q.X. Guan

300. Hyphoderma nudicephalum Gilb. & M. Blackw.

301. Hyphoderma pinicola Yurchenko & Sheng H. Wu

302. Hyphoderma setigerum (Fr.) Donk

303. Hyphoderma subsetigerum Sheng H. Wu

304. Hyphoderma tenuissimum C.L. Zhao & Q.X. Guan

305. Hyphodermella laevigata Yue Li & S.H. He *

306. Hyphodermella rosae (Bres.) Nakasone

307. Hyphodontia pallidula (Bres.) J. Erikss.

308. Hypholoma fasciculare (Huds.) P. Kumm.

309. Hypholoma lateritium (Schaeff.) P. Kumm.

310. Inocutis mikadoi (Lloyd) Y.C. Dai & F. Wu

311. Inocutis rheades (Pers.) Fiasson & Niemelä

312. Inonotus griseus L.W. Zhou

313. Inonotus tenuicontextus L.W. Zhou & W.M. Qin *

314. Irpex alboflavescens Yue Li et al. *

315. Irpex consors Berk.

316. Irpex laceratus (N. Maek. et al.) C.C. Chen & Sheng H. Wu

317. Irpex lacteus (Fr.) Fr.

318. Irpex rosettiformis C.C. Chen & Sheng H. Wu

319. Irpex subulatus (Ryvarden) Z.B. Liu & Y.C. Dai

320. Junghuhnia crustacea (Jungh.) Ryvarden

321. Junghuhnia nitida (Pers.) Ryvarden

322. Junghuhnia pseudominuta H.S. Yuan & Y.C. Dai

323. Kneiffiella abdita Riebesehl & Langer

324. Kneiffiella serpentiformis (Langer) Riebesehl & Langer

325. Kuehneromyces mutabilis (Schaeff.) Singer & A.H. Sm.

326. Laetiporus cremeiporus Y. Ota & T. Hatt.

327. Laetiporus versisporus (Lloyd) Imazeki

328. Laxitextum bicolor (Pers.) Lentz

329. Lentinula edodes (Berk.) Pegler

330. Lentinus arcularius (Batsch) Zmitr.

331. Lentinus brumalis (Pers.) Zmitr.

332. Lentinus connatus Berk.

333. Lentinus fuscus Lloyd

334. Lentinus squarrosulus Mont.

335. Lentinus tigrinus (Bull.) Fr.

336. Lenzites vespaceus (Pers.) Ryvarden

337. Leptoporus mollis (Pers.) Quél.

338. Leucoinocybe auricoma (Har. Takah.) Matheny

339. Lopharia cinerascens (Schwein.) G. Cunn.

340. Lopharia mirabilis (Berk. & Broome) Pat.

341. Luteoporia tenuissima K.Y. Luo et al. *

342. Lyomyces bambusinus C.L. Zhao

343. Lyomyces cremeus C.L. Zhao

344. Lyomyces crustosus (Pers.) P. Karst.

345. Lyomyces daweishanensis J.H. Dong & C.L. Zhao

346. Lyomyces mascarensis Riebesehl et al.

347. Lyomyces microfasciculatus (Yurchenko & Sheng H. Wu) Riebesehl & Langer

348. Lyomyces ochraceoalbus C.L. Zhao

349. Lyomyces orientalis Riebesehl, Yurch. & Langer

350. Lyomyces punctatomarginatus Qi Li & C.L. Zhao

351. Lyomyces vietnamensis (Yurchenko & Sheng H. Wu) Riebesehl & Langer

352. Lyomyces yunnanensis C.L. Zhao

353. Macrosporia nanlingensis (B.K. Cui & C.L. Zhao) B.K. Cui & Xing Ji

354. Marasmiellus albofuscus (Berk. & M.A. Curtis) Singer

355. Marasmiellus alneus Singer

356. Marasmiellus candidus (Fr.) Singer

357. Marasmiellus corticum Singer

358. Marasmiellus dendroegrus Singer

359. Marasmiellus nivosus (Berk.) Singer

360. Marasmiellus sprucei (Berk.) Singer

361. Marasmius albostipitatus Chun Y. Deng & T.H. Li

362. Marasmius aurantiacus I. Hino

363. Marasmius aurantioferrugineus Hongo

364. Marasmius bambusiniformis Singer

365. Marasmius bambusinus Fr.

366. Marasmius bellus Berk.

367. Marasmius berteroi (Lév.) Murrill

368. Marasmius brunneoaurantiacus Antonín & Buyck

369. Marasmius chordalis Fr.

370. Marasmius cohaerens (Pers.) Cooke & Quél.

371. Marasmius coklatus Desjardin, Retn. & E. Horak

372. Marasmius epiphyllus (Pers.) Fr.

373. Marasmius ferrugineus Berk. & Broome

374. Marasmius floriceps Berk. & M.A. Curtis

375. Marasmius graminicola Speg.

376. Marasmius grandiviridis Wannathes et al.

377. Marasmius haematocephalus (Mont.) Fr.

378. Marasmius hinnuleus Berk. & M.A. Curtis

379. Marasmius hymeniicephalus (Sacc.) Singer

380. Marasmius jinfoshanensis Chun Y. Deng & Gafforov

381. Marasmius maximus Hongo

382. Marasmius nigrobrunneus (Pat.) Sacc.

383. Marasmius nigrodiscus (Peck) Halling

384. Marasmius occultatiformis Antonín et al.

385. Marasmius pellucidus Berk. & Broome

386. Marasmius plicatulus Peck

387. Marasmius pulcherripes Peck

388. Marasmius purpureostriatus Hongo

389. Marasmius rotalis Berk. & Broome

390. Marasmius ruber Singer

391. Marasmius ruforotula Singer

392. Marasmius siccus (Schwein.) Fr.

393. Marasmius subabundans Chun Y. Deng & T.H. Li

394. Marasmius suthepensis Wannathes et al.

395. Marasmius wisteriae Antonín et al.

396. Mariorajchenbergia subcavernulosa (Y.C. Dai & Sheng H. Wu) Gibertoni & C.R.S. Lira

397. Megacollybia platyphylla (Pers.) Kotl. & Pouzar

398. Megasporoporia bannaensis B.K. Cui & Hai J. Li

399. Megasporoporia setulosa (Henn.) Rajchenb.

400. Mensularia rhododendri F. Wu et al. *

401. Meripilus eminens (Y.C. Dai) Rajchenb. & Westph.

402. Meripilus giganteus (Pers.) P. Karst.

403. Meripilus lineatus (Pers.) Westph. & Rajchenb.

404. Meripilus pouzarii (Vampola & Vlasák) Westph. & Rajchenb.

405. Meripilus rhododendri (Y.C. Dai et al.) Westph. & Rajchenb. *

406. Meripilus vitreus (Pers.) Rajchenb. & Westph

407. Meruliopsis leptocystidiata C.C. Chen & Sheng H. Wu

408. Meruliopsis rhizomorpha Y.C. Dai et al. *

409. Metuloidea fragrans (A. David & Tortič) Miettinen

410. Metuloidea murashkinskyi (Burt) Miettinen & Spirin

411. Microporus affinis (Blume & T. Nees) Kuntze

412. Microporus xanthopus (Fr.) Kuntze

413. Mucidula mucida (Schrad.) Pat.

414. Mycena abramsii (Murrill) Murrill

415. Mycena acicula (Schaeff.) P. Kumm.

416. Mycena adnexa T. Bau & Q. Na

417. Mycena albiceps (Peck) Gilliam

418. Mycena alcalina (Fr.) P. Kumm.

419. Mycena alphitophora (Berk.) Sacc.

420. Mycena aurantiomarginata (Fr.) Quél.

421. Mycena capillaripes Peck

422. Mycena citricolor (Berk. & M.A. Curtis) Sacc.

423. Mycena corynephora Maas Geest.

424. Mycena epipterygia (Scop.) Gray

425. Mycena filopes (Bull.) P. Kumm.

426. Mycena galericulata (Scop.) Gray

427. Mycena galopus (Pers.) P. Kumm.

428. Mycena haematopus (Pers.) P. Kumm.

429. Mycena laevigata Gillet

430. Mycena leaiana (Berk.) Sacc.

431. Mycena leptocephala (Pers.) Gillet

432. Mycena maculata P. Karst.

433. Mycena metata (Fr.) P. Kumm.

434. Mycena pearsoniana Dennis ex Singer

435. Mycena polygramma (Bull.) Gray,

436. Mycena pura (Pers.) P. Kumm.

437. Mycena robusta (A.H. Sm.) Maas Geest.

438. Mycena rosea Gramberg

439. Mycena rosella (Fr.) P. Kumm.

440. Mycena seminau A.L.C. Chew & Desjardin

441. Mycena semivestipes (Peck) A.H. Sm.

442. Mycena stylobates (Pers.) P. Kumm.

443. Mycena zephirus (Fr.) P. Kumm.

444. Mycetinis salalis (Desjardin & Redhead) Redhead

445. Mycoacia livida (Pers.) Zmitr.

446. Mycoacia neotuberculata S.H. He et al.

447. Mycoaciella bispora (Stalpers) J. Erikss. & Ryvarden

448. Myxarium cinnamomescens (Raitv.) Raitv.

449. Myxomphalia maura (Fr.) Hora

450. Naematelia aurantialba (Bandoni & M. Zang) Millanes & Wedin

451. Naematoloma gracile Hongo

452. Neoantrodiella gypsea (Yasuda) Y.C. Dai et al.

453. Neodatronia sinensis B.K. Cui et al.

454. Neofavolus alveolaris (DC.) Sotome & T. Hatt.

455. Neofavolus mikawae (Lloyd) Sotome & T. Hatt.

456. Neofomitella guangxiensis B.K. Cui & Xing Ji

457. Neolentinus lepideus (Fr.) Redhead & Ginns

458. Neomensularia kanehirae (Yasuda) F. Wu et al.

459. Nidula niveotomentosa (Henn.) Lloyd

460. Nigroporus vinosus (Berk.) Murrill

461. Obba rivulosa (Berk. & M.A. Curtis) Miettinen & Rajchenb.

462. Osmoporus mexicanus (Mont.) Y.C. Dai & S.H. He

463. Pallidohirschioporus biformis (Fr.) Y.C. Dai et al.

464. Pallidohirschioporus brastagii (Corner) Y.C. Dai et al.

465. Panellus bambusicola Q.Y. Zhang & Y.C. Dai

466. Panellus microspermus Q.Y. Zhang et al. *

467. Panellus pubescens Z.S. Bi

468. Panellus pusillus (Pers. ex Lév.) Burds. & O.K. Mill.

469. Panellus stipticus (Bull.) P. Karst.

470. Panellus yunnanensis Q.Y. Zhang & Y.C. Dai

471. Panus conchatus (Bull.) Fr.

472. Panus neostrigosus Drechsler-Santos & Wartchow

473. Panus rudis Fr.

474. Peniophora bicornis Hjortstam & Ryvarden

475. Peniophora crassitunicata Boidin, Lanq. & Gilles

476. Peniophora major Y.L. Xu et al. *

477. Peniophora manshurica Parmasto

478. Peniophora ovalispora Boidin, Lanq. & Gilles

479. Peniophora versiformis (Berk. & M.A. Curtis) Bourdot & Galzin

480. Peniophorella crystallifera Yurchenko et al.

481. Peniophorella praetermissa (P. Karst.) K.H. Larss.

482. Peniophorella rude (Bres.) K.H. Larss.

483. Perenniporia fergusii Gilb. & Ryvarden

484. Perenniporia medulla-panis (Jacq.) Donk

485. Perenniporia prunicola Y.C. Dai et al. *

486. Perenniporiopsis minutissima (Yasuda) C.L. Zhao

487. Phaeophlebiopsis himalayensis (Dhingra) Zmitr.

488. Phallus dongsun T.H. Li et al. *

489. Phanerochaete calotricha (P. Karst.) J. Erikss. & Ryvarden

490. Phanerochaete concrescens Spirin & Volobuev

491. Phanerochaete leptocystidiata Y.L. Xu & S.H. He

492. Phanerochaete rhizomorpha C.C. Chen et al.

493. Phanerochaete sanguineocarnosa Floudas & Hibbett

494. Phanerochaete sordida (P. Karst.) J. Erikss. & Ryvarden

495. Phanerochaete stereoides Sheng H. Wu

496. Phanerochaetella formosana C.C. Chen & Sheng H. Wu

497. Phanerodontia chrysosporium (Burds.) Hjortstam & Ryvarden

498. Phellinus igniarius (L.) Quél.

499. Phellinus monticola L.W. Zhou & Y.C. Dai

500. Phellinus piceicola B.K. Cui & Y.C. Dai

501. Phlebia acerina Peck

502. Phlebia lilascens (Bourdot) J. Erikss. & Hjortstam

503. Phlebia radiata Fr.

504. Phlebia tremellosa (Schrad.) Nakasone & Burds.

505. Phlebiopsis alba (Sang H. Lin & Z.C. Chen) C.C. Chen & Sheng H. Wu

506. Phlebiopsis castanea (Lloyd) Miettinen & Spirin

507. Phlebiopsis crassa (Lév.) Floudas & Hibbett

508. Phlebiopsis gigantea (Fr.) Jülich

509. Phlebiopsis odontoidea C.C. Chen & Sheng H. Wu

510. Phlebiporia bubalina Jia J. Chen et al.

511. Pholiota abietis A.H. Sm. & Hesler

512. Pholiota adiposa (Batsch) P. Kumm.

513. Pholiota flammans (Batsch) P. Kumm.

514. Pholiota fusa (Batsch) Singer

515. Pholiota limonella (Peck) Sacc.

516. Pholiota multicingulata E. Horak

517. Pholiota spumosa (Fr.) Singer

518. Pholiota squarrosa (Vahl) P. Kumm.

519. Phylloporia nandinae L.W. Zhou & Y.C. Dai

520. Phylloporia radiata L.W. Zhou *

521. Phylloporia tiliae L.W. Zhou

522. Phylloporus rubiginosus M.A. Neves & Halling

523. Physalacria bambusae Höhn.

524. Picipes ailaoshanensis B.K. Cui et al.

525. Picipes badius (Pers.) Zmitr. & Kovalenko

526. Picipes dictyopus (Mont.) B.K. Cui et al.

527. Pleurotus citrinopileatus Singer

528. Pleurotus cornucopiae (Paulet) Quél.

529. Pleurotus djamor (Rumph. ex Fr.) Boedijn

530. Pleurotus eryngii (DC.) Quél.

531. Pleurotus floridanus Singer

532. Pleurotus ostreatus (Jacq.) P. Kumm.

533. Pleurotus pulmonarius (Fr.) Quél.

534. Plicaturopsis crispa (Pers.) D.A. Reid

535. Pluteus albidus Beeli

536. Pluteus atricapillus (Batsch) Fayod

537. Pluteus cervinus (Schaeff.) P. Kumm.

538. Pluteus chrysaegis (Berk. & Broome) Petch

539. Pluteus chrysophlebius (Berk. & M.A. Curtis) Sacc.

540. Pluteus cinereofuscus J.E. Lange

541. Pluteus eos Justo & E.F. Malysheva

542. Pluteus griseodiscus Jiang Xu & T.H. Li

543. Pluteus leoninus (Schaeff.) P. Kumm.

544. Pluteus longistriatus (Peck) Peck

545. Pluteus luteus (Redhead & B. Liu) Redhead

546. Pluteus nanus (Pers.) P. Kumm.

547. Pluteus pouzarianus Singer

548. Pluteus purpureofuscus Jiang Xu et al.

549. Pluteus semibulbosus (Lasch) Quél.

550. Pluteus umbrosus (Pers.) P. Kumm.

551. Podofomes mollis (Sommerf.) Gorjón

552. Podoscypha fulvonitens (Berk.) D.A. Reid

553. Polyporus brevibasidiosus H. Lee et al.

554. Polyporus gracilisporus (H. Lee et al.) Bernicchia & Gorjón

555. Polyporus leprieurii Mont.

556. Polyporus mori (Pollini) Fr.

557. Polyporus rugulosus Lév.

558. Polyporus tuberaster (Jacq. ex Pers.) Fr.

559. Poriella subacida (Peck) C.L. Zhao

560. Porogramme albocincta (Cooke & Massee) Gibertoni

561. Porogramme epimiltina (Berk. & Broome) Y.C. Dai et al.

562. Porotheleum fimbriatum (Pers.) Fr.

563. Postia hirsuta L.L. Shen & B.K. Cui

564. Postia tephroleuca (Fr.) Jülich

565. Prunulus diosmus (Krieglst. & Schwöbel) C. Hahn

566. Psathyrella conica T. Bau & J.Q. Yan

567. Psathyrella oboensis Desjardin & B.A. Perry

568. Pseudofavolus tenuis (Fr.) G. Cunn.

569. Pseudohydnum brunneiceps Y.L. Chen et al.

570. Pseudohydnum gelatinosum (Scop.) P. Karst.

571. Pseudolagarobasidium subvinosum (Berk. & Broome) Sheng H. Wu

572. Pseudomarasmius pallidocephalus (Gilliam) R.H. Petersen

573. Pseudomerulius aureus (Fr.) Jülich

574. Pseudomerulius curtisii (Berk.) Redhead & Ginns

575. Pusillomyces asetosus (Antonín et al.) J.S. Oliveira

576. Radulomyces confluens (Fr.) M.P. Christ.

577. Sidera lenis (Pat.) Hjortstam & Spooner

578. Radulomyces hydnoides J.H. Dong & C.L. Zhao

579. Ramaria botrytis (Pers.) Bourdot

580. Ramaria gracilis (Pers.) Quél.

581. Rectipilus fasciculatus (Pers.) Agerer

582. Resinicium bicolor (Alb. & Schwein.) Parmasto

583. Resinomycena mirabilis (Singer) Redhead & Nagas.

584. Resupinatus alboniger (Pat.) Singer

585. Resupinatus applicatus (Batsch) Gray

586. Resupinatus trichotis (Pers.) Singer

587. Rhizochaete radicata (Henn.) Gresl. et al.

588. Rhizochaete subradicata Yue Li & S.H. He

589. Rhizochaete variegata Q.Y. Zhang et al. *

590. Rhizomarasmius cunninghamietorum Chun Y. Deng et al. *

591. Rhodofomes subfeei (B.K. Cui & M.L. Han) B.K. Cui et al.

592. Rigidoporus cirratus (Hjortstam & Ryvarden) L.W. Zhou

593. Rigidoporus crocatus (Pat.) Ryvarden

594. Rigidoporus cuneatus (Murrill) F. Wu et al.

595. Rigidoporus ginkgonis (Y.C. Dai) F. Wu et al.

596. Rigidoporus lineatus (Pers.) Ryvarden

597. Rigidoporus microporus (Sw.) Overeem

598. Rigidoporus populinus (Schumach.) Pouzar

599. Rigidoporus pouzarii Vampola & Vlasák

600. Roridomyces roridus (Fr.) Rexer

601. Ryvardenia campyla (Berk.) Rajchenb.

602. Sanghuangporus baumii (Pilát) L.W. Zhou & Y.C. Dai

603. Sanghuangporus quercicola Lin Zhu & B.K. Cui

604. Sanghuangporus sanghuang (Sheng H. Wu et al.) Sheng H. Wu et al.

605. Sanghuangporus weigelae (T. Hatt. & Sheng H. Wu) Sheng H. Wu et al.

606. Sanguinoderma rude (Berk.) Y.F. Sun et al.

607. Sanguinoderma rugosum (Blume & T. Nees) Y.F. Sun et al.

608. Sarcoporia longitubulata Vlasák & Spirin

609. Schizophyllum commune Fr.

610. Scopuloides ellipsoidea S.H. He et al.

611. Scopuloides hydnoides (Cooke & Massee) Hjortstam & Ryvarden

612. Scytinostroma acystidiatum Q.Y. Zhang et al. *

613. Scytinostroma renisporum Boidin et al.

614. Scytinostroma duriusculum (Berk. & Broome) Donk

615. Scytinostroma macrospermum Q.Y. Zhang et al.

616. Scytinostroma odoratum (Fr.) Donk

617. Scytinostroma renisporum Boidin et al.

618. Scytinostroma subduriusculum Yue Li et al.

619. Scytinostroma subrenisporum Yue Li et al. *

620. Sidera tibetica Z.B. Liu et al.

621. Sidera vulgaris (Fr.) Miettinen

622. Sirobasidium magnum Boedijn

623. Sistotrema brinkmannii (Bres.) J. Erikss.

624. Skeletocutis albocremea A. David

625. Skeletocutis alutacea (J. Lowe) Jean Keller

626. Skeletocutis amorpha (Fr.) Kotl. & Pouzar

627. Skeletocutis biguttulata (Romell) Niemelä

628. Skeletocutis chrysella Niemelä

629. Skeletocutis diluta (Rajchenb.) A. David & Rajchenb.

630. Skeletocutis futilis Miettinen & A. Korhonen

631. Skeletocutis kuehneri A. David

632. Skeletocutis lepida A. Korhonen & Miettinen

633. Skeletocutis liangdongii Z.B. Liu et al. *

634. Skeletocutis mopanshanensis C.L. Zhao

635. Skeletocutis nivea (Jungh.) Jean Keller

636. Skeletocutis pseudo-odora L.F. Fan & Jing Si *

637. Skeletocutis semipileata (Peck) Miettinen & A. Korhonen

638. Skeletocutis sinica Z.B. Liu et al. *

639. Sparassis crispa (Wulfen) Fr.

640. Steccherinum aurantilaetum (Corner) Bernicchia & Gorjón

641. Steccherinum fimbriatum (Pers.) J. Erikss.

642. Steccherinum helvolum (Zipp. ex Lév.) S. Ito

643. Steccherinum ochraceum (Pers. ex J.F. Gmel.) Gray

644. Steccherinum tenuissimum C.L. Zhao & Y.X. Wu

645. Stereum gausapatum (Fr.) Fr.

646. Stereum hirsutum (Willd.) Pers.

647. Stereum lithocarpi Y.C. Dai

648. Stereum ostrea (Blume & T. Nees) Fr.

649. Stereum rugosum Pers.

650. Stereum sanguinolentum (Alb. & Schwein.) Fr.

651. Stereum subtomentosum Pouzar

652. Stropharia lignicola E.J. Tian

653. Subulicystidium acerosum S.H. He & S.L. Liu *

654. Tapinella atrotomentosa (Batsch) Šutara

655. Terana coerulea (Lam.) Kuntze

656. Tetrapyrgos alba (Berk. & M.A. Curtis) E. Horak

657. Tetrapyrgos nigripes (Fr.) E. Horak

658. Tetrapyrgos subcinerea (Berk. & Broome) E. Horak

659. Tomentella bryophila (Pers.) M.J. Larsen

660. Tomentella casiae H.S. Yuan & Y.Q. Zhu*

661. Tomentella ellisii (Sacc.) Jülich & Stalpers

662. Tomentella fuscocinerea (Pers.) Donk

663. Tomentella guiyangensis H.S. Yuan & Y.Q. Zhu*

664. Tomentella olivaceomarginata H.S. Yuan & Y.Q. Zhu*

665. Tomentella rotundata H.S. Yuan & Y.Q. Zhu*

666. Tomentella stuposa (Link) Stalpers

667. Trachydermella tsunodae (Yasuda ex Lloyd) B.K. Cui & Y.F. Sun

668. Trametes betulina (L.) Pilát

669. Trametes cinnabarina (Jacq.) Fr.

670. Trametes coccinea (Fr.) Hai J. Li & S.H. He

671. Trametes elegans (Spreng.) Fr.

672. Trametes gibbosa (Pers.) Fr.

673. Trametes hirsuta (Wulfen) Lloyd

674. Trametes lacerata Lloyd

675. Trametes manilaensis (Lloyd) Teng

676. Trametes membranacea (Sw.) Kreisel

677. Trametes ochracea (Pers.) Gilb. & Ryvarden

678. Trametes orientalis (Yasuda) Imazeki

679. Trametes pubescens (Schumach.) Pilát

680. Trametes strumosa (Fr.) Zmitr., Wasser & Ezhov

681. Trametes suaveolens (L.) Fr.

682. Trametes trogii Berk.

683. Trametes vernicipes (Berk.) Zmitr., Wasser & Ezhov

684. Trametes villosa (Sw.) Kreisel

685. Trametopsis cervina (Schwein.) Tomšovský

686. Trechispora candidissima (Schwein.) Bondartsev & Singer

687. Trechispora constricta S.L. Liu et al.

688. Trechispora gracilis S.L. Liu & L.W. Zhou

689. Trechispora larssonii S.L. Liu et al. *

690. Trechispora latihypha S.L. Liu et al.

691. Trechispora longiramosa S.L. Liu et al. *

692. Trechispora nivea (Pers.) K.H. Larss.

693. Trechispora odontioidea (L.) Lloyd

694. Trechispora sinensis S.L. Liu et al. *

695. Trechispora wenshanensis K.Y. Luo & C.L. Zhao

696. Tremella cerebriformis Chee J. Chen

697. Tremella cinnabarina Bull.

698. Tremella flava Chee J. Chen

699. Tremella foliacea Pers.

700. Tremella fuciformis Berk.

701. Tremella mesenterica (Schaeff.) Pers.

702. Tremella samoensis Lloyd

703. Trichaptum byssogenum (Jungh.) Ryvarden

704. Trichaptum pergamenum (Fr.) G. Cunn.

705. Truncospora ochroleuca (Berk.) Pilát

706. Truncospora tephropora (Mont.) Zmitr.

707. Tubaria punicea (A.H. Sm. & Hesler) Ammirati et al.

708. Tubulicium bambusicola S.H. He & S.L. Liu

709. Tyromyces chioneus (Fr.) P. Karst.

710. Tyromyces galactinus (Berk.) J. Lowe

711. Tyromyces lacteus (Fr.) Murrill

712. Tyromyces kmetii (Bres.) Bondartsev & Singer

713. Vanderbylia fraxinea (Bull.) D.A. Reid

714. Vanderbylia robiniophila (Murrill) B.K. Cui & Y.C. Dai

715. Vararia amphithallica Boidin et al.

716. Vararia athabascensis Gilb.

717. Veluticeps fasciculata Jiao Yang & S.H. He

718. Vizzinia nigripes (Angelini et al.) Ševčíková & Kolařík

719. Vitreoporus dichrous (Fr.) Zmitr.

720. Xenasmatella ardosiaca (Bourdot & Galzin) Stalpers

721. Xeromphalina brevipes T. Bau & L.N. Liu

722. Xeromphalina campanella (Batsch) Kühner & Maire

723. Xeromphalina kauffmanii A.H. Sm.

724. Xeromphalina tenuipes (Schwein.) A.H. Sm.

725. Xanthoperenniporia maackiae (Bondartsev & Ljub.) B.K. Cui & Xing Ji

726. Xanthoperenniporia tenuis (Schwein.) B.K. Cui & Xing Ji

727. Xylobolus princeps (Jungh.) Boidin

728. Xylobolus spectabilis (Klotzsch) Boidin

729. Xylobolus subpileatus (Berk. & M.A. Curtis) Boidin

730. Xylodon exilis Yurchenko et al.

731. Xylodon flaviporus (Berk. & M.A. Curtis ex Cooke) Riebesehl & Langer

732. Xylodon nesporii (Bres.) Hjortstam & Ryvarden

733. Xylodon niemelaei (Sheng H. Wu) Hjortstam & Ryvarden

734. Xylodon nongravis (Lloyd) Che C. Chen & Sheng H. Wu

735. Xylodon ovisporus (Corner) Riebesehl & Langer

736. Xylodon paradoxus (Schrad.) Chevall

737. Xylodon reticulatus (Che C. Chen & Sheng H. Wu) Che C. Chen & Sheng H. Wu

738. Xylodon taiwanianus (Sheng H. Wu) Hjortstam & Ryvarden

2.5 贵州省木生大型担子菌优势类群分析

本研究共记录贵州省木生大型担子菌738种，隶属于担子菌门的3纲16目74科253属。其中，3个纲分别为担子菌纲Agaricomycetes、花耳纲Dacrymycetes和银耳纲Tremellomycetes。担子菌纲为贵州省木生大型担子菌的绝对优势类群，共包含14目71科244属717种，占总物种数的97.15%。相比之下，花耳纲仅包括1目1科6属12种，占总物种数的1.63%；银耳纲包括1目2科3属9种，占总物种数的1.22%，二者在物种数量上均处于绝对次要地位。

在目级水平上，贵州省木生大型担子菌的优势目主要为多孔菌目Polyporales、蘑菇目Agaricales和锈革孔菌目Hymenochaetales。这3个目共计包含50科194属601种，分别占总科数的67.57%、总属数的76.68%和总物种数的81.44% (图 2)。其中，多孔菌目的物种多样性最为丰富，包含22科100属253种，占总物种数的34.28%；其次为蘑菇目，共记录19科63属218种，占比为29.54%；锈革孔菌目包括11科31属130种，占比为17.62%。其余13个目合计包含24科59属137种，物种占比为18.56%。在这13个目中，物种数量相对较多的为红菇目Russulales和木耳目Auriculariales，分别记录有47种和30种。

在科级水平上，贵州省木生大型担子菌中共有10个优势科(图 3)，分别为多孔菌科Polyporaceae (36属92种)、锈革孔菌科Hymenochaetaceae (16属67种)、小菇科Mycenaceae (8属47种)、小皮伞科Marasmiaceae (4属42种)、类脐菇科Omphalotaceae (7属31种)、木耳科Auriculariaceae (8属26种)、原毛平革菌科Phanerochaetaceae (9属23种)、裂孔菌科Schizoporaceae (4属23种)、囊耙齿菌科Irpicaceae (9属20种)以及隔孢伏革菌科Peniophoraceae (5属20种)。上述10个优势科共计包含106属391种，分别占总属数和总物种数的41.90%和52.98%。其余64个科共包含126属324种，分别占总属数的49.80%和总物种数的43.90%。此外，还有21属23种尚未被明确归入已知科级分类单元，分别占总属数和总物种数的8.30%和3.12%。

在属级层面，共识别出11个优势属(图4)，分别为小皮伞属Marasmius (35种)、小菇属Mycena (30种)、刺革菌属Hymenochaete (28种)、栓孔菌属Trametes (17种)、光柄菇属Pluteus(16种)、集毛孔菌属Coltricia (15种)、干皮孔菌属Skeletocutis (15种)、灵芝属Ganoderma (13种)、金钱菌属Collybiopsis (12种)、疏革菌属Lyomyces (11种)和糙孢孔菌属Trechispora (10种)。上述11个优势属共计包含202种，占总物种数的27.37%；其余242个属共包含536种，占总物种数的72.63%。

此外，在本研究确认的738种贵州省木生大型担子菌中，有48个物种以贵州省作为其模式产地或副模式产地，占总物种数的6.50%，进一步彰显了贵州省在木生大型担子菌分类学和物种形成研究方面的重要地位。

3 结论

收起

真菌作为生物圈中第二大生物类群，具有极其丰富的物种多样性以及复杂多样的形态特征和演化历程。目前，全球已知真菌物种数已超过20万种(https://nmdc.cn/fungalnames/，获取时间：2025年12月1日)。其中，木生大型担子菌形态类型复杂多样，涵盖多孔菌、齿菌、革菌、伞菌和珊瑚菌等多个形态类型(Dong et al. 2024；Spirin et al. 2024；Zhao et al. 2025)，具有非常丰富的物种多样性。

近年来，随着生物多样性保护意识的不断增强以及系统分类学方法的广泛应用，我国真菌多样性研究取得了一系列重要进展(戴玉成和庄剑云 2010；王科等2021)。然而，由于研究区域、研究方法和研究类群侧重点的不同，不同地区及不同类群间的研究仍存在明显不均衡现象。其中，贵州省作为中国西南地区的重要生态区域，其木生大型担子菌多样性研究相对滞后。尽管已有研究者对贵州省部分区域的大型真菌多样性开展了探索性调查，如仁怀市大型真菌初步研究(赵本学等 2023)及桴焉自然保护区的相关调查(杨云深等 2025)，但总体而言，尚缺乏针对全省尺度的系统梳理与综合总结。现有关于贵州省木生大型担子菌的研究多零散分布于多篇文献之中，缺乏整合性的物种多样性框架(戴玉成等 2021；赵本学等 2023；Li et al. 2025；Liu et al. 2025a, 2025b；Xu et al. 2025；杨云深等 2025)。此外，贵州省地处云贵高原腹地，其地形、气候和植被类型高度多样，与真菌多样性研究基础较为成熟的云南省相比，仍具有巨大的研究潜力。

本研究首次对贵州省木生大型担子菌进行了系统整理与统计，明确其共计隶属于担子菌门3纲16目74科253属738种。该成果显著丰富了贵州省的真菌物种多样性认知，并为今后开展系统分类、生态功能及资源利用等相关研究提供了重要的基础数据。值得注意的是，在本研究中发现，蘑菇目作为贵州省木生大型担子菌的优势目之一，其物种多样性超过了传统认为占优势地位的锈革孔菌目；在优势科与优势属层面，同样有多个主要类群隶属于蘑菇目的类群，如小菇科Mycenaceae、小菇属Mycena以及小皮伞科Marasmiaceae和小皮伞属Marasmius等。近期研究表明，蘑菇目中包含大量以木材为基质的真菌类群，其生态功能和演化意义长期以来可能被低估(Dong et al. 2025)，相关问题仍有待在未来研究中进一步深入探讨。

此外，本研究特别关注了以贵州省作为模式产地的木生大型担子菌物种。戴玉成等(2021)在研究中国森林大型真菌多样性与系统学时，仅记录贵州省拥有8个模式物种，显著低于其邻近地区，如云南省(314个)和广西壮族自治区(41个)。经过近4年的系统研究，该数字已大幅提升至48个。一方面，这表明贵州省蕴藏着尚未充分揭示的真菌多样性潜力；另一方面，也展示了近年来对贵州省真菌资源的调查与系统分类研究正快速推进。

总体而言，本研究系统补全了贵州省木生大型担子菌多样性的基础资料，明确了其主要优势类群，为后续的资源挖掘、开发与相关生态研究提供了重要的数据支撑。

4 数据可用性声明

收起

本数据集已存储至科学数据银行(ScienceDB)期刊数据平台 https://scidb.cn/jwxb，可被所有用户公开访问，按照CC BY-NC-ND 4.0协议共享和使用。

[数据引用格式]

赵恒, 陈芊, 张欣, 罗凯月，李彪，员瑗，吴芳，何双辉，曾广宇，庄磊，戴玉成，邓春英. 中国贵州省木生大型担子菌编目[DS/OL]. V1. Science Data Bank, 2026[2026-03-16]. https://cstr.cn/31253.11.sciencedb.36968. CSTR:31253.11.sciencedb.36968.

赵恒, 陈芊, 张欣, 罗凯月，李彪，员瑗，吴芳，何双辉，曾广宇，庄磊，戴玉成，邓春英. 中国贵州省木生大型担子菌编目[DS/OL]. V1. Science Data Bank, 2026[2026-03-16]. https://doi.org/10.57760/sciencedb.36968. DOI:10.57760/sciencedb.36968.

基金

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国家自然科学基金(32360008)

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2026年第45卷第4期

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文章信息

doi: 10.13346/j.mycosystema.250358

接收时间：2025-12-15
首发时间：2026-04-30
出版时间：2026-04-22

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出版历史

收稿日期：2025-12-15
录用日期：2026-01-05

基金

National Natural Sciences Foundation of China(32360008)

国家自然科学基金(32360008)

作者信息

¹ 北京林业大学生态与自然保护学院，北京 100083

² 中国科学院沈阳应用生态研究所森林生态与保育重点实验室，辽宁沈阳 110016

³ 重庆交通大学建筑与城市规划学院，重庆 400074

⁴ 广西壮族自治区林业科学研究院，广西南宁 530002

⁵ 哈尔滨市农业科学院，黑龙江哈尔滨 150028

⁶ 贵州科学院贵州省生物研究所，贵州贵阳 550009

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2种不同金属材料的力学参数

科 Family	属数 Number of genus	种数 Number of species	占总种数比例 Percentage of total species (%)	属 Genus	种数 Number of species	占总种数比例 Percentage of total species (%)
鹅膏菌科Amanitaceae	2	11	5.26	鹅膏菌属 Amanita	10	4.78
小菇科 Mycenaceae	2	12	5.74	丝盖伞属 Inocybe	5	2.39
多孔菌科 Polyporaceae	8	14	6.70	蜡蘑属 Laccaria	5	2.39
红菇科 Russulaceae	3	23	11.00	小皮伞属 Marasmius	6	2.87
				小菇属 Mycena	11	5.26
				光柄菇属 Pluteus	5	2.39
				红菇属 Russula	17	8.13
				栓菌属 Trametes	5	2.39

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