Article(id=1322924795272241965, tenantId=1146029695717560320, journalId=1322258771877134531, issueId=1322924673771647833, articleNumber=null, orderNo=null, doi=10.14067/j.cnki.1673-923x.2026.09.017, pmid=null, cstr=null, oa=null, hot=null, price=null, onlineType=0, articleFormat=0, articleType=null, articleTypeStr=null, receivedDate=1763049600000, receivedDateStr=2025-11-14, revisedDate=1766937600000, revisedDateStr=2025-12-29, acceptedDate=null, acceptedDateStr=null, onlineDate=1793339449946, onlineDateStr=2026-10-30, pubDate=1790265600000, pubDateStr=2026-09-25, doiRegisterDate=null, doiRegisterDateStr=null, onlineIssueDate=1793339449946, onlineIssueDateStr=2026-10-30, onlineJustAcceptDate=null, onlineJustAcceptDateStr=null, onlineFirstDate=null, onlineFirstDateStr=null, sourceXml=null, magXml=null, createTime=1793339449946, creator=13701087609, updateTime=1793339449946, updator=13701087609, issue=Issue{id=1322924673771647833, tenantId=1146029695717560320, journalId=1322258771877134531, year='2026', volume='46', issue='9', pageStart='1', pageEnd='250', issueExtLink='null', onlineDate='null', pubDate='1790265600000', pubDateStr='2026-09-25', beforeIssueId=null, nextIssueId=null, price=null, status=1, issueComplete=1, articleOrder=1, issueType=-1, specialIssue=null, createTime=1793339420978, creator='13701087609', updateTime=1793344623946, updator='13701087609', preIssue=null, nextIssue=null, articleTotal=null, ext={EN=IssueExt(id=1322946497460961720, tenantId=1146029695717560320, journalId=1322258771877134531, issueId=1322924673771647833, language=EN, specialIssueTitle=, coverIllustrator=null, specialIssueEditor=, specialIssueAbout=), CN=IssueExt(id=1322946497460961721, tenantId=1146029695717560320, journalId=1322258771877134531, issueId=1322924673771647833, language=CN, specialIssueTitle=, coverIllustrator=null, specialIssueEditor=, specialIssueAbout=)}, issueFiles=null, downloadFileDto=null}, startPage=191, endPage=200, ext={EN=ArticleExt(id=1322924795599397678, articleId=1322924795272241965, tenantId=1146029695717560320, journalId=1322258771877134531, language=EN, title=Effect of stand density on understory plant diversity and soil nutrients in Juglans mandshurica natural secondary forest, columnId=null, journalTitle=Journal of Central South University of Forestry & Technology, columnName=null, runingTitle=null, highlight=null, articleAbstract=
【Objective】

To investigate the effects of stand densities on the structure of understory plant communities, species diversity and soil nutrient content in Juglans mandshurica forests, and to reveal the intrinsic relationship between plant diversity and soil nutrients.

【Method】

This study focused on natural secondary forests of Juglans mandshurica on Mao'er mountain with different densities (840, 680 and 480 trees·hm-2). Through field community surveys and indoor soil analysis, the species composition, importance values, diversity index of understory woody and herbaceous plants were investigated, and soil nutrient indicators at different soil layer depths were measured for correlation analysis.

【Result】

1) A total of 65 plant species were surveyed under the Juglans mandshurica tree forest throughout the study area, including 16 woody species belonging to 11 families and 14 genera, and 49 herbaceous species belonging to 28 families and 45 genera; 2) The dominant species of the woody layer is Syringa reticulata subsp. amurensis, with a maximum importance value of 85.23% at a stand density of 480 trees·hm-2; the dominant species of the herbaceous layer is Beckmannia syzigachne, with a maximum importance value of 50.49% at a stand density of 680 trees·hm-2; 3) Species richness is highest when the stand density is 680 trees·hm-2, with the richness indices of woody and herbaceous plants being 11 and 31, respectively; the Simpson and Shannon-Wiener diversity indices show a trend of first increasing and then decreasing as stand density decreases, reaching their highest values at a stand density of 680 trees·hm-2; the evenness index increases as stand density decreases, and the highest evenness index for herbaceous plants is 0.81 at a stand density of 480 trees·hm-2; 4) The nutrient content in the soil is highest in the 0-10 cm layer. When the stand density is 680 trees·hm-2, the contents of organic matter, total nitrogen, alkaline nitrogen and available phosphorus in this soil layer are significantly higher than those in other density treatments. Woody plant diversity is extremely significantly positively correlated with total soil nitrogen content and significantly negatively correlated with soil pH, while herbaceous plant richness is extremely significantly positively correlated with the content of available phosphorus.

【Conclusion】

A stand density of 680 trees·hm-2 is the optimal density in the study area for maintaining understory plant diversity in Juglans mandshurica forests and enhancing soil fertility. Stand density indirectly regulates the composition and diversity of plant communities by altering the soil environment, this study will provide a reasonable management density for secondary forests of Juglans mandshurica and offer a scientific basis for the sustainable management of Juglans mandshurica.

, authors=Yutong Wang, Yuedong Lyu, Jianying Zhang, Wenhui Kong, Zhongling Liu, Wenzhi Ge, authorsList=Yutong Wang, Yuedong Lyu, Jianying Zhang, Wenhui Kong, Zhongling Liu, Wenzhi Ge, authorCompany=null, correspAuthors=Zhongling Liu, Wenzhi Ge, authorNote=null, correspAuthorsNote=null, copyrightStatement=© 2026 Journal of Central South University of Forestry & Technology, copyrightOwner=Journal of Central South University of Forestry & Technology, extLink=null, articleAbsUrl=null, sourceXml=null, magXml=null, pdfUrl=null, pdf=null, pdfFileSize=null, pdfExtLink=null, richHtmlUrl=null, mobilePdfUrl=null, reviewReport=null, pdfFirstPage=null, abstractGraph=null, abstractGraphContent=null, abstractVideo=null, citation=null, cebUrl=null, magXmlContent=null, mapNumber=null, fund=null), CN=ArticleExt(id=1322924795947524911, articleId=1322924795272241965, tenantId=1146029695717560320, journalId=1322258771877134531, language=CN, title=林分密度对胡桃楸天然次生林林下植物多样性与土壤养分的影响, columnId=1322924188427760267, journalTitle=中南林业科技大学学报, columnName=生态学, runingTitle=null, highlight=null, articleAbstract=
【目的】

探究林分密度对胡桃楸Juglans mandshurica林下植物群落结构、物种多样性及土壤养分含量的影响,揭示植物多样性与土壤养分的内在联系。

【方法】

以帽儿山不同林分密度(840、680、480株·hm-2)的胡桃楸天然次生林为研究对象,采用野外群落调查与室内土壤理化分析相结合的方法,测定林下植物的物种组成、重要值、多样性指数以及不同土层土壤养分指标,并进行相关性分析。

【结果】

1)在整个研究区域内胡桃楸林下共调查到65种植物。其中木本植物16种,隶属于11科14属;草本植物49种,隶属于28科45属。2)木本层的优势种为暴马丁香Syringa reticulata subsp. amurensis,其重要值在480株·hm-2的林分密度中最大,为85.23%;草本层的优势种为菵草Beckmannia syzigachne,其重要值在680株·hm-2的林分密度中最大,为50.49%。3)物种丰富度在林分密度为680株·hm-2时最高,其木本植物和草本植物的丰富度指数分别为11和31;Simpson、Shannon-Wiener多样性指数随着林分密度的减小表现为先升高后降低的趋势,林分密度为680株·hm-2时多样性指数值最高;均匀度指数随着林分密度的减小而增大,当林分密度为480株·hm-2时草本植物的均匀度指数最高,为0.81。4)土壤养分含量在0~10 cm土层最高,当林分密度为680株·hm-2时,该土层中的有机质、全氮、碱解氮和有效磷含量显著高于其他林分密度。木本植物多样性与土壤全氮含量呈极显著正相关,与土壤pH值呈显著负相关;草本植物丰富度与有效磷含量呈极显著正相关。

【结论】

680株·hm-2是本研究区域内维持胡桃楸林下植物多样性和提升土壤肥力的最适林分密度。

, authors=王雨童, 吕跃东, 张建瑛, 孔文汇, 刘忠玲, 葛文志, authorsList=王雨童, 吕跃东, 张建瑛, 孔文汇, 刘忠玲, 葛文志, authorCompany=null, correspAuthors=刘忠玲, 葛文志, authorNote=

王雨童(),实习研究员,硕士。

, correspAuthorsNote=
刘忠玲(),研究员,博士
葛文志(),研究员,硕士。
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林分密度对胡桃楸天然次生林林下植物多样性与土壤养分的影响
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王雨童 , 吕跃东 , 张建瑛 , 孔文汇 , 刘忠玲 , 葛文志
中南林业科技大学学报 | 生态学 2026,46(9): 191-200
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中南林业科技大学学报 |生态学 2026 , 46 (9) : 191 -200
林分密度对胡桃楸天然次生林林下植物多样性与土壤养分的影响
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王雨童 , 吕跃东, 张建瑛, 孔文汇, 刘忠玲 , 葛文志
作者信息
  • 黑龙江省林业科学研究所 黑龙江省速生林木培育重点实验室,黑龙江 哈尔滨 150081
通讯作者:
刘忠玲(),研究员,博士
葛文志(),研究员,硕士。
作者简介:

王雨童(),实习研究员,硕士。

Effect of stand density on understory plant diversity and soil nutrients in Juglans mandshurica natural secondary forest
Yutong Wang , Yuedong Lyu, Jianying Zhang, Wenhui Kong, Zhongling Liu , Wenzhi Ge
Affiliations
  • Heilongjiang Forestry Institute, Key Laboratory of Fast-growing Tree Cultivating of Heilongjiang Province, Harbin 150081, Heilongjiang, China
出版时间: 2026-09-25 doi: 10.14067/j.cnki.1673-923x.2026.09.017
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【目的】

探究林分密度对胡桃楸Juglans mandshurica林下植物群落结构、物种多样性及土壤养分含量的影响,揭示植物多样性与土壤养分的内在联系。

【方法】

以帽儿山不同林分密度(840、680、480株·hm-2)的胡桃楸天然次生林为研究对象,采用野外群落调查与室内土壤理化分析相结合的方法,测定林下植物的物种组成、重要值、多样性指数以及不同土层土壤养分指标,并进行相关性分析。

【结果】

1)在整个研究区域内胡桃楸林下共调查到65种植物。其中木本植物16种,隶属于11科14属;草本植物49种,隶属于28科45属。2)木本层的优势种为暴马丁香Syringa reticulata subsp. amurensis,其重要值在480株·hm-2的林分密度中最大,为85.23%;草本层的优势种为菵草Beckmannia syzigachne,其重要值在680株·hm-2的林分密度中最大,为50.49%。3)物种丰富度在林分密度为680株·hm-2时最高,其木本植物和草本植物的丰富度指数分别为11和31;Simpson、Shannon-Wiener多样性指数随着林分密度的减小表现为先升高后降低的趋势,林分密度为680株·hm-2时多样性指数值最高;均匀度指数随着林分密度的减小而增大,当林分密度为480株·hm-2时草本植物的均匀度指数最高,为0.81。4)土壤养分含量在0~10 cm土层最高,当林分密度为680株·hm-2时,该土层中的有机质、全氮、碱解氮和有效磷含量显著高于其他林分密度。木本植物多样性与土壤全氮含量呈极显著正相关,与土壤pH值呈显著负相关;草本植物丰富度与有效磷含量呈极显著正相关。

【结论】

680株·hm-2是本研究区域内维持胡桃楸林下植物多样性和提升土壤肥力的最适林分密度。

胡桃楸天然次生林  /  林分密度  /  植物多样性  /  土壤养分
【Objective】

To investigate the effects of stand densities on the structure of understory plant communities, species diversity and soil nutrient content in Juglans mandshurica forests, and to reveal the intrinsic relationship between plant diversity and soil nutrients.

【Method】

This study focused on natural secondary forests of Juglans mandshurica on Mao'er mountain with different densities (840, 680 and 480 trees·hm-2). Through field community surveys and indoor soil analysis, the species composition, importance values, diversity index of understory woody and herbaceous plants were investigated, and soil nutrient indicators at different soil layer depths were measured for correlation analysis.

【Result】

1) A total of 65 plant species were surveyed under the Juglans mandshurica tree forest throughout the study area, including 16 woody species belonging to 11 families and 14 genera, and 49 herbaceous species belonging to 28 families and 45 genera; 2) The dominant species of the woody layer is Syringa reticulata subsp. amurensis, with a maximum importance value of 85.23% at a stand density of 480 trees·hm-2; the dominant species of the herbaceous layer is Beckmannia syzigachne, with a maximum importance value of 50.49% at a stand density of 680 trees·hm-2; 3) Species richness is highest when the stand density is 680 trees·hm-2, with the richness indices of woody and herbaceous plants being 11 and 31, respectively; the Simpson and Shannon-Wiener diversity indices show a trend of first increasing and then decreasing as stand density decreases, reaching their highest values at a stand density of 680 trees·hm-2; the evenness index increases as stand density decreases, and the highest evenness index for herbaceous plants is 0.81 at a stand density of 480 trees·hm-2; 4) The nutrient content in the soil is highest in the 0-10 cm layer. When the stand density is 680 trees·hm-2, the contents of organic matter, total nitrogen, alkaline nitrogen and available phosphorus in this soil layer are significantly higher than those in other density treatments. Woody plant diversity is extremely significantly positively correlated with total soil nitrogen content and significantly negatively correlated with soil pH, while herbaceous plant richness is extremely significantly positively correlated with the content of available phosphorus.

【Conclusion】

A stand density of 680 trees·hm-2 is the optimal density in the study area for maintaining understory plant diversity in Juglans mandshurica forests and enhancing soil fertility. Stand density indirectly regulates the composition and diversity of plant communities by altering the soil environment, this study will provide a reasonable management density for secondary forests of Juglans mandshurica and offer a scientific basis for the sustainable management of Juglans mandshurica.

Juglans mandshurica  /  stand densities  /  plant diversity  /  soil nutrients
王雨童, 吕跃东, 张建瑛, 孔文汇, 刘忠玲, 葛文志. 林分密度对胡桃楸天然次生林林下植物多样性与土壤养分的影响. 中南林业科技大学学报, 2026 , 46 (9) : 191 -200 . DOI: 10.14067/j.cnki.1673-923x.2026.09.017
Yutong Wang, Yuedong Lyu, Jianying Zhang, Wenhui Kong, Zhongling Liu, Wenzhi Ge. Effect of stand density on understory plant diversity and soil nutrients in Juglans mandshurica natural secondary forest[J]. Journal of Central South University of Forestry & Technology, 2026 , 46 (9) : 191 -200 . DOI: 10.14067/j.cnki.1673-923x.2026.09.017
  • 黑龙江省省属科研院所科研业务费项目(JC-202506)
  • 国家重点研发计划项目(2017YFD0600605)
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2026年第46卷第9期
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doi: 10.14067/j.cnki.1673-923x.2026.09.017
  • 接收时间:2025-11-14
  • 首发时间:2026-10-30
  • 出版时间:2026-09-25
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  • 收稿日期:2025-11-14
  • 修回日期:2025-12-29
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黑龙江省省属科研院所科研业务费项目(JC-202506)
国家重点研发计划项目(2017YFD0600605)
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    黑龙江省林业科学研究所 黑龙江省速生林木培育重点实验室,黑龙江 哈尔滨 150081

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刘忠玲(),研究员,博士
葛文志(),研究员,硕士。
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2种不同金属材料的力学参数
科
Family
属数
Number of
genus
种数
Number of
species
占总种数比例
Percentage of
total species (%)
属
Genus
种数
Number of
species
占总种数比例
Percentage of total
species (%)
鹅膏菌科Amanitaceae 2 11 5.26 鹅膏菌属 Amanita 10 4.78
小菇科 Mycenaceae 2 12 5.74 丝盖伞属 Inocybe 5 2.39
多孔菌科 Polyporaceae 8 14 6.70 蜡蘑属 Laccaria 5 2.39
红菇科 Russulaceae 3 23 11.00 小皮伞属 Marasmius 6 2.87
小菇属 Mycena 11 5.26
光柄菇属 Pluteus 5 2.39
红菇属 Russula 17 8.13
栓菌属 Trametes 5 2.39
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