Article(id=1322927260776399513, tenantId=1146029695717560320, journalId=1322258771877134531, issueId=1322924673771647833, articleNumber=null, orderNo=null, doi=10.14067/j.cnki.1673-923x.2026.09.005, pmid=null, cstr=null, oa=null, hot=null, price=null, onlineType=0, articleFormat=0, articleType=null, articleTypeStr=null, receivedDate=1756224000000, receivedDateStr=2025-08-27, revisedDate=1762358400000, revisedDateStr=2025-11-06, acceptedDate=null, acceptedDateStr=null, onlineDate=1793340037768, onlineDateStr=2026-10-30, pubDate=1790265600000, pubDateStr=2026-09-25, doiRegisterDate=null, doiRegisterDateStr=null, onlineIssueDate=1793340037768, onlineIssueDateStr=2026-10-30, onlineJustAcceptDate=null, onlineJustAcceptDateStr=null, onlineFirstDate=null, onlineFirstDateStr=null, sourceXml=null, magXml=null, createTime=1793340037768, creator=13701087609, updateTime=1793340037768, updator=13701087609, issue=Issue{id=1322924673771647833, tenantId=1146029695717560320, journalId=1322258771877134531, year='2026', volume='46', issue='9', pageStart='1', pageEnd='250', issueExtLink='null', onlineDate='null', pubDate='1790265600000', pubDateStr='2026-09-25', beforeIssueId=null, nextIssueId=null, price=null, status=1, issueComplete=1, articleOrder=1, issueType=-1, specialIssue=null, createTime=1793339420978, creator='13701087609', updateTime=1793344623946, updator='13701087609', preIssue=null, nextIssue=null, articleTotal=null, ext={EN=IssueExt(id=1322946497460961720, tenantId=1146029695717560320, journalId=1322258771877134531, issueId=1322924673771647833, language=EN, specialIssueTitle=, coverIllustrator=null, specialIssueEditor=, specialIssueAbout=), CN=IssueExt(id=1322946497460961721, tenantId=1146029695717560320, journalId=1322258771877134531, issueId=1322924673771647833, language=CN, specialIssueTitle=, coverIllustrator=null, specialIssueEditor=, specialIssueAbout=)}, issueFiles=null, downloadFileDto=null}, startPage=51, endPage=61, ext={EN=ArticleExt(id=1322927261187441306, articleId=1322927260776399513, tenantId=1146029695717560320, journalId=1322258771877134531, language=EN, title=Influence of climate and neighborhood competition on radial growth of Pinus koraiensis in eastern Heilongjiang mountainous areas, columnId=null, journalTitle=Journal of Central South University of Forestry & Technology, columnName=null, runingTitle=null, highlight=null, articleAbstract=
【Objective】

The study aims to investigate the effects of climatic factors and interspecific competition on the radial growth of Pinus koraiensis, and to provide scientific support for predicting forest productivity and carbon sink potential under future climate change, as well as the conservation and sustainable management of regional forest resources.

【Method】

Pinus koraiensis trees from two national nature reserves (Xiaobeihu and Xilinhe) in the mountainous eastern Heilongjiang Province were selected as research materials. Dendrochronological techniques were adopted to acquire radial growth data. Combined with meteorological datasets and neighborhood competition indices, correlation analysis and linear mixed-effects models (LMM) were applied to quantify the independent impacts of climate and competition on the radial growth of Pinus koraiensis. Variance decomposition was further performed to calculate their relative contributions, so as to explore the effects of climatic conditions and neighborhood competition on radial growth as well as corresponding regional disparities.

【Result】

1) The radial growth of Pinus koraiensis in the Xiaobeihu study area showed significantly positive correlations with temperatures of the previous winter and current spring. Linear mixed-effects model (LMM) outputs indicated that the diameter at breast height (DBH) of competing trees achieved the optimal fitting performance with cumulative radial growth over the past 20 years; 2) Radial growth of Pinus koraiensis in the Xilinhe study area was primarily controlled by March temperature of the current year. LMM results revealed that DBH of neighboring competitive trees had the best fitting effect with cumulative growth over the recent 10 years; 3) The contribution proportions of climatic factors and inter-tree competition differed along the latitudinal gradient. With increasing latitude, the relative contribution rate of climate rose from 39% to 88%, while that of competition decreased from 61% to 11%.

【Conclusion】

This study indicates that the dominant factor regulating radial growth of Pinus koraiensis shifts from biotic competition to climatic limitation along mountainous eastern Heilongjiang. Thinning can alleviate competitive pressure in low-latitude forest stands, while high-latitude forests should be alert to growth stress induced by spring warming.

, authors=Haoxia Zhang1, Luyao Wang1, Wenjun Zhang2, Cunyong Ju1, authorsList=Haoxia Zhang, Luyao Wang, Wenjun Zhang, Cunyong Ju, authorCompany=null, correspAuthors=Cunyong Ju, authorNote=null, correspAuthorsNote=null, copyrightStatement=© 2026 Journal of Central South University of Forestry & Technology, copyrightOwner=Journal of Central South University of Forestry & Technology, extLink=null, articleAbsUrl=null, sourceXml=null, magXml=null, pdfUrl=null, pdf=null, pdfFileSize=null, pdfExtLink=null, richHtmlUrl=null, mobilePdfUrl=null, reviewReport=null, pdfFirstPage=null, abstractGraph=null, abstractGraphContent=null, abstractVideo=null, citation=null, cebUrl=null, magXmlContent=null, mapNumber=null, fund=null), CN=ArticleExt(id=1322927261711729307, articleId=1322927260776399513, tenantId=1146029695717560320, journalId=1322258771877134531, language=CN, title=黑龙江东部山区气候和邻体竞争对红松径向生长的影响, columnId=1322924153761837409, journalTitle=中南林业科技大学学报, columnName=林学, runingTitle=null, highlight=null, articleAbstract=
【目的】

研究气候因子和种间竞争对红松Pinus koraiensis径向生长的影响,为未来气候变化背景下森林生产力与碳汇潜力的预测、地区森林资源保护和可持续经营提供科学依据。

【方法】

以黑龙江东部山区小北湖和细鳞河2个国家级自然保护区的红松为研究对象,利用树木年轮学方法获取径向生长数据,结合气象数据和邻体竞争指标,采用相关分析和线性混合效应模型(Linear mixed-effects model,LMM)方法量化气候与竞争对红松径向生长的影响,并通过方差分解解析各因子的相对贡献,探究气候因子与邻体竞争对红松径向生长的影响与区域差异。

【结果】

1)小北湖研究区红松径向生长与前一年冬季及当年春季气温呈显著正相关。线性混合效应模型结果显示,红松竞争树胸径与过去20 a的累积生长量之间的拟合效果最优。2)细鳞河研究区红松主要受当年3月气温的影响。线性混合效应模型结果显示,竞争树胸径与过去10 a的累积生长量之间的拟合效果最佳。3)两地竞争与气候的贡献度存在纬度差异,随着纬度的升高,气候贡献率由39%增至88%,而竞争贡献率由61%降至11%。

【结论】

黑龙江东部山区红松径向生长的影响因子由生物竞争转为气候限制,低纬度林区可通过疏伐缓解竞争,高纬度林区需警惕春季升温带来的生长胁迫。

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张豪霞(),硕士研究生。

, correspAuthorsNote=
琚存勇(),副教授,博士。
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Intraspecific competition in a Cunninghamia lanceolata plantation with different age and diameter classes[J]. Journal of Zhejiang A & F University, 2015, 32(3): 353-360., articleTitle=Intraspecific competition in a Cunninghamia lanceolata plantation with different age and diameter classes, refAbstract=null)], funds=[Fund(id=1322927269165007550, tenantId=1146029695717560320, journalId=1322258771877134531, articleId=1322927260776399513, awardId=32371860, language=CN, fundingSource=国家自然科学基金项目(32371860), fundOrder=null, country=null)], companyList=[AuthorCompany(id=1322927262068245148, tenantId=1146029695717560320, journalId=1322258771877134531, articleId=1322927260776399513, xref=1., ext=[AuthorCompanyExt(id=1322927262076633757, tenantId=1146029695717560320, journalId=1322258771877134531, articleId=1322927260776399513, companyId=1322927262068245148, language=EN, country=null, province=null, city=null, postcode=null, companyName=null, departmentName=null, remark=1.a. 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黑龙江东部山区气候和邻体竞争对红松径向生长的影响
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张豪霞 1 , 王露瑶 1 , 张文军 2 , 琚存勇 1
中南林业科技大学学报 | 林学 2026,46(9): 51-61
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中南林业科技大学学报 |林学 2026 , 46 (9) : 51 -61
黑龙江东部山区气候和邻体竞争对红松径向生长的影响
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张豪霞1 , 王露瑶1, 张文军2, 琚存勇1
作者信息
  • 1.东北林业大学 a. 林学院;b. 森林生态系统可持续经营教育部重点实验室,黑龙江 哈尔滨 150040
  • 2.黑龙江细鳞河国家级自然保护区管理局,黑龙江 鹤岗 154101
通讯作者:
琚存勇(),副教授,博士。
作者简介:

张豪霞(),硕士研究生。

Influence of climate and neighborhood competition on radial growth of Pinus koraiensis in eastern Heilongjiang mountainous areas
Haoxia Zhang1 , Luyao Wang1, Wenjun Zhang2, Cunyong Ju1
Affiliations
  • 1.a. College of Forestry; b. Key Laboratory of Sustainable Forest Ecosystem Management, Ministry of Education, Northeast Forestry University, Harbin 150040, Heilongjiang, China
  • 2.Administration of Xilinhe National Nature Reserve, Hegang 154101, Heilongjiang, China
出版时间: 2026-09-25 doi: 10.14067/j.cnki.1673-923x.2026.09.005
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【目的】

研究气候因子和种间竞争对红松Pinus koraiensis径向生长的影响,为未来气候变化背景下森林生产力与碳汇潜力的预测、地区森林资源保护和可持续经营提供科学依据。

【方法】

以黑龙江东部山区小北湖和细鳞河2个国家级自然保护区的红松为研究对象,利用树木年轮学方法获取径向生长数据,结合气象数据和邻体竞争指标,采用相关分析和线性混合效应模型(Linear mixed-effects model,LMM)方法量化气候与竞争对红松径向生长的影响,并通过方差分解解析各因子的相对贡献,探究气候因子与邻体竞争对红松径向生长的影响与区域差异。

【结果】

1)小北湖研究区红松径向生长与前一年冬季及当年春季气温呈显著正相关。线性混合效应模型结果显示,红松竞争树胸径与过去20 a的累积生长量之间的拟合效果最优。2)细鳞河研究区红松主要受当年3月气温的影响。线性混合效应模型结果显示,竞争树胸径与过去10 a的累积生长量之间的拟合效果最佳。3)两地竞争与气候的贡献度存在纬度差异,随着纬度的升高,气候贡献率由39%增至88%,而竞争贡献率由61%降至11%。

【结论】

黑龙江东部山区红松径向生长的影响因子由生物竞争转为气候限制,低纬度林区可通过疏伐缓解竞争,高纬度林区需警惕春季升温带来的生长胁迫。

红松  /  径向生长  /  邻体竞争  /  气候响应
【Objective】

The study aims to investigate the effects of climatic factors and interspecific competition on the radial growth of Pinus koraiensis, and to provide scientific support for predicting forest productivity and carbon sink potential under future climate change, as well as the conservation and sustainable management of regional forest resources.

【Method】

Pinus koraiensis trees from two national nature reserves (Xiaobeihu and Xilinhe) in the mountainous eastern Heilongjiang Province were selected as research materials. Dendrochronological techniques were adopted to acquire radial growth data. Combined with meteorological datasets and neighborhood competition indices, correlation analysis and linear mixed-effects models (LMM) were applied to quantify the independent impacts of climate and competition on the radial growth of Pinus koraiensis. Variance decomposition was further performed to calculate their relative contributions, so as to explore the effects of climatic conditions and neighborhood competition on radial growth as well as corresponding regional disparities.

【Result】

1) The radial growth of Pinus koraiensis in the Xiaobeihu study area showed significantly positive correlations with temperatures of the previous winter and current spring. Linear mixed-effects model (LMM) outputs indicated that the diameter at breast height (DBH) of competing trees achieved the optimal fitting performance with cumulative radial growth over the past 20 years; 2) Radial growth of Pinus koraiensis in the Xilinhe study area was primarily controlled by March temperature of the current year. LMM results revealed that DBH of neighboring competitive trees had the best fitting effect with cumulative growth over the recent 10 years; 3) The contribution proportions of climatic factors and inter-tree competition differed along the latitudinal gradient. With increasing latitude, the relative contribution rate of climate rose from 39% to 88%, while that of competition decreased from 61% to 11%.

【Conclusion】

This study indicates that the dominant factor regulating radial growth of Pinus koraiensis shifts from biotic competition to climatic limitation along mountainous eastern Heilongjiang. Thinning can alleviate competitive pressure in low-latitude forest stands, while high-latitude forests should be alert to growth stress induced by spring warming.

Pinus koraiensis  /  radial growth  /  neighboring tree competition  /  climate response
张豪霞, 王露瑶, 张文军, 琚存勇. 黑龙江东部山区气候和邻体竞争对红松径向生长的影响. 中南林业科技大学学报, 2026 , 46 (9) : 51 -61 . DOI: 10.14067/j.cnki.1673-923x.2026.09.005
Haoxia Zhang, Luyao Wang, Wenjun Zhang, Cunyong Ju. Influence of climate and neighborhood competition on radial growth of Pinus koraiensis in eastern Heilongjiang mountainous areas[J]. Journal of Central South University of Forestry & Technology, 2026 , 46 (9) : 51 -61 . DOI: 10.14067/j.cnki.1673-923x.2026.09.005
  • 国家自然科学基金项目(32371860)
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2026年第46卷第9期
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doi: 10.14067/j.cnki.1673-923x.2026.09.005
  • 接收时间:2025-08-27
  • 首发时间:2026-10-30
  • 出版时间:2026-09-25
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  • 收稿日期:2025-08-27
  • 修回日期:2025-11-06
基金
国家自然科学基金项目(32371860)
作者信息
    1.东北林业大学 a. 林学院;b. 森林生态系统可持续经营教育部重点实验室,黑龙江 哈尔滨 150040
    2.黑龙江细鳞河国家级自然保护区管理局,黑龙江 鹤岗 154101

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琚存勇(),副教授,博士。
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2种不同金属材料的力学参数
科
Family
属数
Number of
genus
种数
Number of
species
占总种数比例
Percentage of
total species (%)
属
Genus
种数
Number of
species
占总种数比例
Percentage of total
species (%)
鹅膏菌科Amanitaceae 2 11 5.26 鹅膏菌属 Amanita 10 4.78
小菇科 Mycenaceae 2 12 5.74 丝盖伞属 Inocybe 5 2.39
多孔菌科 Polyporaceae 8 14 6.70 蜡蘑属 Laccaria 5 2.39
红菇科 Russulaceae 3 23 11.00 小皮伞属 Marasmius 6 2.87
小菇属 Mycena 11 5.26
光柄菇属 Pluteus 5 2.39
红菇属 Russula 17 8.13
栓菌属 Trametes 5 2.39
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