Article(id=1309812603862483095, tenantId=1146029695717560320, journalId=1308397449582313577, issueId=1309812542579503925, articleNumber=null, orderNo=null, doi=10.7605/gdlxb.2025.078, pmid=null, cstr=null, oa=null, hot=null, price=null, onlineType=0, articleFormat=0, articleType=null, articleTypeStr=research-article, receivedDate=1744214400000, receivedDateStr=2025-04-10, revisedDate=1745510400000, revisedDateStr=2025-04-25, acceptedDate=null, acceptedDateStr=null, onlineDate=1790213259901, onlineDateStr=2026-09-24, pubDate=1774972800000, pubDateStr=2026-04-01, doiRegisterDate=null, doiRegisterDateStr=null, onlineIssueDate=1790213259901, onlineIssueDateStr=2026-09-24, onlineJustAcceptDate=null, onlineJustAcceptDateStr=null, onlineFirstDate=null, onlineFirstDateStr=null, sourceXml=null, magXml=null, createTime=1790213259901, creator=13701087609, updateTime=1790213259901, updator=13701087609, issue=Issue{id=1309812542579503925, tenantId=1146029695717560320, journalId=1308397449582313577, year='2026', volume='28', issue='2', pageStart='429', pageEnd='842', issueExtLink='null', onlineDate='null', pubDate='1774972800000', pubDateStr='2026-04-01', beforeIssueId=null, nextIssueId=null, price=null, status=1, issueComplete=1, articleOrder=1, issueType=-1, specialIssue=null, createTime=1790213245291, creator='13701087609', updateTime=1790213366052, updator='13701087609', preIssue=null, nextIssue=null, articleTotal=null, ext={EN=IssueExt(id=1309813049133981868, tenantId=1146029695717560320, journalId=1308397449582313577, issueId=1309812542579503925, language=EN, specialIssueTitle=, coverIllustrator=null, specialIssueEditor=, specialIssueAbout=), CN=IssueExt(id=1309813049133981869, tenantId=1146029695717560320, journalId=1308397449582313577, issueId=1309812542579503925, language=CN, specialIssueTitle=, coverIllustrator=null, specialIssueEditor=, specialIssueAbout=)}, issueFiles=null, downloadFileDto=null}, startPage=708, endPage=723, ext={EN=ArticleExt(id=1309812604147695768, articleId=1309812603862483095, tenantId=1146029695717560320, journalId=1308397449582313577, language=EN, title=The Silurian-Devonian radiation of vascular plants and the potential effects on terrestrial carbon stocks, columnId=1309812571910267828, journalTitle=Journal of Palaeogeography (Chinese Edition), columnName=BIOPALAEOGEOGRAPHY AND PALAEOECOLOGY, runingTitle=null, highlight=null, articleAbstract=
Vascular plants,as the major clade of land plants,represent the most important plant group that colonized the land during the Paleozoic. Fossil records demonstrate that vascular plants first appeared in the Silurian and then underwent significant radiation during the Devonian. Plants from the Silurian-Devonian periods represent the ancestors or stem groups of modern lineages such as the lycopsids,equisetopsids,filicopsids,and seed plants,foreshadowing their earliest differentiation. The total genus-level diversity of vascular plants gradually increased from the Silurian,reached the first peak during the Pragian,which was then followed by two decline-rise fluctuations,with one peak registrated during the late Givetian and another peak during the late Famennian. The vegetation types of the Silurian to Early Devonian include rhyniopsid herblands,zosterophyllopsid herblands and zosterophyllopsid-early euphyllophyte mixed herblands,with sporadic occurrences of shrubs and wetlands. During the Middle and Late Devonian,herblands,wetlands and shrubs continued to develop,while emerged new vegetation types such as cladoxylopsid forests,lycopsid forests,and progymnosperm forests. Along with the Silurian-Devonian radiation of vascular plants,the vegetation carbon stock significantly expanded,and the soil inorganic carbon stock also increased,as indicated by the dramatic increase of calcareous paleosols. Some Lower Devonian paleosols have inorganic carbon densities comparable to the modern maximum values. Since the Middle Devonian,the records of coal(paleo-histosols)gradually increased,indicating a significant rise in the amount of soil organic carbon. The increase of terrestrial carbon burial during the Silurian-Devonian was indicated by the accumulation of carbon-rich soils such as histosols and calcareous soils. Through diagenesis,carbon from surface system was converted into paleosols as a part of the lithosphere carbon stock,with an effect of long-term carbon sequestration. Future efforts,on one side,should be put into the systematics and classification of plant fossils,and as well as the study of paleobotanical community ecology and the classification of early vegetation. On the other side,the reconstruction of whole-plants and their communities,model simulations,and other methods could be conducted to estimate the vegetation carbon stocks of different time intervals. The information of surface soil carbon storage of different time intervals and burial efficiency of paleosols should be used to constrain the carbon cycle processes in Earth System models.
, authors=Jinzhuang XUE
1, Tao ZHONG
1, Hui WANG
1, Bingxin LI
1, Qiqi YANG
1, Jiashu WANG
2, authorsList=Jinzhuang XUE, Tao ZHONG, Hui WANG, Bingxin LI, Qiqi YANG, Jiashu WANG, authorCompany=null, correspAuthors=null, authorNote=
XUE Jinzhuang,born in 1981,is an associate professor of Peking University. His research focuses on plant terrestrialization and its effects on Earth system. E-mail: pkuxue@pku.edu.cn.
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*, columnId=1309812572090622902, journalTitle=古地理学报, columnName=生物古地理学及古生态, runingTitle=null, highlight=null, articleAbstract=
维管植物是陆地植物中最具优势的一大分支,为古生代植物登陆演化的主体。化石记录表明,维管植物最早出现于志留纪,在泥盆纪迎来辐射发展,而志留纪—泥盆纪植物是现代支系如石松类、木贼类、真蕨类及种子植物的祖先类群(干群),预示着这些现代支系的分化。维管植物大化石的属级总体多样性自志留纪逐步上升,至早泥盆世布拉格期达到第1次峰值,之后呈现2次下降、上升波动,在吉维特期晚期及法门期晚期分别达到峰值。志留纪—早泥盆世植被包括莱尼蕨类草地、工蕨类草地、工蕨类—早期真叶植物混生草地等,零星出现灌丛、湿地;中、晚泥盆世草地、湿地、灌丛继续发育,新出现枝蕨类森林、石松类森林和前裸子植物森林等。随着志留纪—泥盆纪植物的辐射演化,植被碳库显著扩大,钙质古土壤记录剧增,表明土壤无机碳库也呈扩大趋势,一些下泥盆统古土壤中的无机碳密度已与现代最高值相近;中泥盆世以来煤(古有机土)的记录逐渐增多,表明土壤有机碳含量的显著增大;因此,志留纪—泥盆纪陆地碳埋藏的增加主要体现在有机土、钙质土等富碳土壤记录的增多。通过成岩作用,古土壤将表层系统中的碳转变为岩石圈碳库的一部分,进而达到长期碳封存的效果。未来需继续加强植物化石的系统分类及古植物群落生态、早期植被分类和区划的研究,进一步通过古植物个体及群落重建、模型模拟等研究估算当时的植被碳库,定量估算不同地质时期的地表土壤碳储量以及古土壤碳的埋藏效率,以约束地球系统模型中的碳循环过程。
, authors=薛进庄
1, 钟涛
1, 王徽
1, 李炳鑫
1, 杨琦琦
1, 王嘉树
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1, address=1 The Key Laboratory of Orogenic Belts and Crustal Evolution,School of Earth and Space Sciences, Peking University,Beijing 100871,China, bio={"content":"
XUE Jinzhuang,born in 1981,is an associate professor of Peking University. His research focuses on plant terrestrialization and its effects on Earth system. E-mail: pkuxue@pku.edu.cn.
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薛进庄,男,1981年生,北京大学地球与空间科学学院副教授,主要从事植物登陆演化及其环境效应的研究。E-mail: pkuxue@pku.edu.cn。
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薛进庄,男,1981年生,北京大学地球与空间科学学院副教授,主要从事植物登陆演化及其环境效应的研究。E-mail: pkuxue@pku.edu.cn。
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2008. 龙潭楔叶的再研究: 兼论华南晚泥盆世楔叶目的演化.
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2009. 中国志留纪陆生植物研究综述.
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2010. 华南古生代陆生维管植物多样性演变.
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2021. 华南板块古生代维管植物数据集.
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2010. 古生物学术语冠群和干群的释义问题.
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14: 602., articleTitle=Fern-like plants establishing the understory of the Late Devonian Xinhang lycopsid forest, refAbstract=null)], funds=[Fund(id=1309812611164766447, tenantId=1146029695717560320, journalId=1308397449582313577, articleId=1309812603862483095, awardId=2022YFF0800200, language=EN, fundingSource=National Key Research and Development Program of China(2022YFF0800200), fundOrder=null, country=null), Fund(id=1309812611231875312, tenantId=1146029695717560320, journalId=1308397449582313577, articleId=1309812603862483095, awardId=2022YFF0800200, language=CN, fundingSource=* 国家重点研发计划项目(2022YFF0800200), fundOrder=null, country=null), Fund(id=1309812611332538609, tenantId=1146029695717560320, journalId=1308397449582313577, articleId=1309812603862483095, awardId=41722201, language=EN, fundingSource=National Natural Science Foundation of China(41722201), fundOrder=null, country=null), Fund(id=1309812611403841779, tenantId=1146029695717560320, journalId=1308397449582313577, articleId=1309812603862483095, awardId=41722201, language=CN, fundingSource=国家自然科学基金项目(41722201), fundOrder=null, country=null)], companyList=[AuthorCompany(id=1309812606295179432, tenantId=1146029695717560320, journalId=1308397449582313577, articleId=1309812603862483095, xref=null, ext=[AuthorCompanyExt(id=1309812606303568041, tenantId=1146029695717560320, journalId=1308397449582313577, articleId=1309812603862483095, companyId=1309812606295179432, language=EN, country=null, province=null, city=null, postcode=null, companyName=null, departmentName=null, remark=1 The Key Laboratory of Orogenic Belts and Crustal Evolution,School of Earth and Space Sciences, Peking University,Beijing 100871,China), AuthorCompanyExt(id=1309812606311956650, tenantId=1146029695717560320, journalId=1308397449582313577, articleId=1309812603862483095, companyId=1309812606295179432, language=CN, country=null, province=null, city=null, postcode=null, companyName=null, departmentName=null, remark=1 北京大学地球与空间科学学院,造山带与地壳演化教育部重点实验室,北京 100871)]), AuthorCompany(id=1309812606450368683, tenantId=1146029695717560320, journalId=1308397449582313577, articleId=1309812603862483095, xref=null, ext=[AuthorCompanyExt(id=1309812606462951596, tenantId=1146029695717560320, journalId=1308397449582313577, articleId=1309812603862483095, companyId=1309812606450368683, language=EN, country=null, province=null, city=null, postcode=null, companyName=null, departmentName=null, remark=2 Laboratory of Geo-specimens Study and Testing,Geological Museum of China,Beijing 100034, China), AuthorCompanyExt(id=1309812606471340205, tenantId=1146029695717560320, journalId=1308397449582313577, articleId=1309812603862483095, companyId=1309812606450368683, language=CN, country=null, province=null, city=null, postcode=null, companyName=null, departmentName=null, remark=2 中国地质博物馆地质标本研究与检测实验室,北京 100034)])], figs=[ArticleFig(id=1309812609738703070, tenantId=1146029695717560320, journalId=1308397449582313577, articleId=1309812603862483095, language=EN, label=Fig.1, caption=
Evolution of large clades of vascular plants through geological time(modified from Cleal and Cascales-Miñana,2014), figureFileSmall=BJAn43gB25B8Q/4+JIbqRA==, figureFileBig=1KXwmOFkTO6YYECNM1vreQ==, tableContent=null), ArticleFig(id=1309812609830977759, tenantId=1146029695717560320, journalId=1308397449582313577, articleId=1309812603862483095, language=CN, label=图 1, caption=
地史时期维管植物大类群的延限及多样性演化(据Cleal and Cascales-Miñana,2014;有修改), figureFileSmall=BJAn43gB25B8Q/4+JIbqRA==, figureFileBig=1KXwmOFkTO6YYECNM1vreQ==, tableContent=null), ArticleFig(id=1309812610028110048, tenantId=1146029695717560320, journalId=1308397449582313577, articleId=1309812603862483095, language=EN, label=Fig.2, caption=
Genus-level diversity of Silurian-Devonian vascular plants (modified from Capel et al., 2022), figureFileSmall=PUifGnrkgPtDuuUFTdILNg==, figureFileBig=3nHIzGfjqRCI/Nf5GNqpmw==, tableContent=null), ArticleFig(id=1309812610103607521, tenantId=1146029695717560320, journalId=1308397449582313577, articleId=1309812603862483095, language=CN, label=图 2, caption=
志留纪—泥盆纪维管植物属级多样性变化(据Capel et al., 2022;有修改) 地质年代符号: Wen,温洛克世; Lud,罗德洛世; Pri,普里道利世; Lock,洛赫考夫期; Pra,布拉格期; Ems,埃姆斯期; Eif,艾菲尔期; Giv,吉维特期; Frs,弗拉期; Fam,法门期。总体多样性、界限穿越者多样性、平均持续多样性的定义参见Capel等(2022)
, figureFileSmall=PUifGnrkgPtDuuUFTdILNg==, figureFileBig=3nHIzGfjqRCI/Nf5GNqpmw==, tableContent=null), ArticleFig(id=1309812610195882210, tenantId=1146029695717560320, journalId=1308397449582313577, articleId=1309812603862483095, language=EN, label=Fig.3, caption=
Reconstruction of several representative Devonian vegetation types, figureFileSmall=LU3o5jj0303GYhl8CRTIog==, figureFileBig=OmKOClFNEMAPgG1uhszBag==, tableContent=null), ArticleFig(id=1309812610267185379, tenantId=1146029695717560320, journalId=1308397449582313577, articleId=1309812603862483095, language=CN, label=图 3, caption=
泥盆纪几类典型植被复原图 A—贵州都匀下泥盆统蟒山群中的工蕨类草地(引自Huang et al., 2025;谭超绘图);B—云南曲靖下泥盆统徐家冲组中的镰蕨类草地(引自Xue et al., 2016的附件材料; 邓珍珍绘图);C—最早的森林——德国Lindlar中泥盆统艾菲尔阶中的枝蕨类森林(引自Berry,2019;Peter Giesen原创);D—华南上泥盆统五通组中的植被景观,包括前裸子植物Archaeopteris森林(后方坡地)、石松类Sublepidodendron森林 (前右)及楔叶类Xihuphyllum草地(前左)(引自Xue et al., 2018;邓珍珍绘图)
, figureFileSmall=LU3o5jj0303GYhl8CRTIog==, figureFileBig=OmKOClFNEMAPgG1uhszBag==, tableContent=null), ArticleFig(id=1309812610338488548, tenantId=1146029695717560320, journalId=1308397449582313577, articleId=1309812603862483095, language=EN, label=Fig.4, caption=
The global carbon cycle of present-day Earth(modified from Friedlingstein et al., 2025;vegetation and soil carbon stocks based on other sources,referring to the main text), figureFileSmall=aiP4+jzrvQQfFGsa3d2Keg==, figureFileBig=8z8I0psX14dl5ixEZxemRA==, tableContent=null), ArticleFig(id=1309812610426568933, tenantId=1146029695717560320, journalId=1308397449582313577, articleId=1309812603862483095, language=CN, label=图 4, caption=
现代全球碳循环示意图(修改自Friedlingstein et al., 2025;植被和土壤碳库据其他资料,见正文), figureFileSmall=aiP4+jzrvQQfFGsa3d2Keg==, figureFileBig=8z8I0psX14dl5ixEZxemRA==, tableContent=null), ArticleFig(id=1309812610497872102, tenantId=1146029695717560320, journalId=1308397449582313577, articleId=1309812603862483095, language=EN, label=Fig.5, caption=
Global patterns of vegetation biomass and NPP of three intervals during the Silurian-Devonian periods, estimated by the SCION-FLORA coupling model(after Gurung et al.,2024), figureFileSmall=CI1MkjepAPaOJNiNpRym2Q==, figureFileBig=ivrDgrbb+vOX35cq6rLAyg==, tableContent=null), ArticleFig(id=1309812610577563879, tenantId=1146029695717560320, journalId=1308397449582313577, articleId=1309812603862483095, language=CN, label=图 5, caption=
SCION-FLORA耦合模型估算的志留纪—泥盆纪3个时期的植被生物量及净初级生产力格局(据Gurung et al.,2024), figureFileSmall=CI1MkjepAPaOJNiNpRym2Q==, figureFileBig=ivrDgrbb+vOX35cq6rLAyg==, tableContent=null), ArticleFig(id=1309812610653061352, tenantId=1146029695717560320, journalId=1308397449582313577, articleId=1309812603862483095, language=EN, label=Fig.6, caption=
Soil carbon processes in arid and humid environments, figureFileSmall=PBjyH/sMpK2CM7c7HehWPA==, figureFileBig=gtUz/JvkFpOA5h+poftLVw==, tableContent=null), ArticleFig(id=1309812610732753129, tenantId=1146029695717560320, journalId=1308397449582313577, articleId=1309812603862483095, language=CN, label=图 6, caption=
干旱和湿润环境中的土壤碳过程 A、B—现代干旱、湿润环境中的土壤无机碳、有机碳过程(修改自Cotrufo and Lavallee(2025));C—云南曲靖下泥盆统桂家屯组中的钙质土(纵向叠覆的2个古土壤序列,干旱环境);D—美国纽约中泥盆世富含土壤有机碳的森林土表面(湿润环境),示直立树干印痕(被水充填的2个凹坑)和样方(2.5 m(纵向)×1.5 m(横向));E—D图森林中挖掘出的树干化石。D、E图片由William E. Stein提供
, figureFileSmall=PBjyH/sMpK2CM7c7HehWPA==, figureFileBig=gtUz/JvkFpOA5h+poftLVw==, tableContent=null), ArticleFig(id=1309812610799861994, tenantId=1146029695717560320, journalId=1308397449582313577, articleId=1309812603862483095, language=EN, label=Fig.7, caption=
Workflow used to study and interpret the Silurian-Devonian vegetation and soils, figureFileSmall=NpzLeE9/Gb40t35gwKY63A==, figureFileBig=dU81UmOrQXF1+IsJCM6ueA==, tableContent=null), ArticleFig(id=1309812610887942380, tenantId=1146029695717560320, journalId=1308397449582313577, articleId=1309812603862483095, language=CN, label=图 7, caption=
志留纪—泥盆纪植被—土壤系统的研究和解释框架示意图 A—古地理底图(据Scotese,2021);B—地球系统模型图(据Matthaeus et al., 2023);C—工蕨类植物化石(据Hao et al., 2010);D—具根系古土壤(据Xue et al., 2016);E—楔叶类植物景观复原(据Huang et al., 2017;邓珍珍绘图)
, figureFileSmall=NpzLeE9/Gb40t35gwKY63A==, figureFileBig=dU81UmOrQXF1+IsJCM6ueA==, tableContent=null), ArticleFig(id=1309812610959245549, tenantId=1146029695717560320, journalId=1308397449582313577, articleId=1309812603862483095, language=EN, label=Table1, caption=
Several representative vegetation types of the Silurian-Devonian periods
, figureFileSmall=null, figureFileBig=null, tableContent=
| 植被型组 | 植被类型 | 产地、层位及植被组成 | 可能的生物量信息 |
| 草地 | 莱尼蕨类草地 | 英国Anglo-Welsh盆地罗德洛统Pterinea层,具多种Cooksonia(Edwards and Richardson,2004) | Gerrienne等(2006)发现1枚近完整保存的Cooksonia标本,显示其植株高33 mm,茎轴宽度0.6~1.0 mm,多次分枝 |
| 莱尼蕨类草地 | 英国Anglo-Welsh盆地下泥盆统Ditton组,具多种Cooksonia、Salopella、Tortilicaulis等(Edwards and Richardson,2004) | 如上 |
| 工蕨类草地 | 加拿大极区Bathurst岛罗德洛统,具cf. Bathurstia sp.、Zosterophyllum sp.等(Kotyk et al., 2002) | Hao等(2010)发现1枚近完整保存的Zosterophyllum标本,显示植株地上部分高98 mm,根系统约20 mm,生物量根茎比约为0.028 |
| 工蕨类草地 | 贵州都匀下泥盆统蟒山群,具Zosterophyllum australianum、Z. baoyangense,混生Adoketophyton、Hedeia等(Huang et al., 2025) | 如上 |
| 工蕨类—早期真叶植物
混生草地 | 云南文山下泥盆统坡松冲组,工蕨纲物种占比43%,早期真叶植物物种占比22%,混生其他种类(Hao and Xue,2013) | 据估算,工蕨类Discalis植株高约20~30 cm,始叶蕨Eophyllophyton植株高约50~80 cm(Hao and Beck,1993;Hao and Xue,2013) |
| 工蕨类—镰蕨类
混生草地 | 云南曲靖下泥盆统徐家冲组,Zosterophyllum和Drepanophycus极为常见,同时混生Hedeia、Huia等(Wang et al., 2002;Xue et al., 2016) | 据Schweitzer(1980),Drepanophycus植株地上部分高50~60 cm;据Xue等(2016),Drepanophycus具复杂、庞大的地下茎系统 |
| 原始鳞木类草地 | 云南曲靖中泥盆统海口组,一些层位具密集、近原位的原始鳞木类植物Minarodendron(郑高峰等,2026) | 郑高峰等(2026)报道Minarodendron茎干最长可达1 m,宽4~5 mm,可能匍匐生长 |
| 森林 | 枝蕨类森林 | 美国纽约州中泥盆统Moscow组,森林主体为Eospermatopteris树木,兼具无脉树目、石松类分子(Stein et al., 2012) | Eospermatopteris树木基部宽50~150 cm,高至少8 m,侧枝逐渐脱落,仅树冠留存;在1200 m2的原位森林样方中,约具200个Eospermatopteris树以及其他植物种类(Stein et al., 2012) |
| 石松类森林 | 安徽广德上泥盆统五通组,森林主体为Guangdedendron树木(“新杭森林”),兼具早期似真蕨类等(Wang et al., 2019;Yang et al., 2024) | “新杭森林”为泥盆纪出露面积最大(至少250 000 m2)的森林,主体为Guangdedendron树木,估计高1.1~7.7 m,但大多低于3.2 m高,密度达38株/m2(Wang et al., 2019) |
| 前裸子植物森林 | Archaeopteris/Callixylon在晚泥盆世全球广布;美国纽约州中泥盆统Plattekill组,发现较为罕见的Archaeopteris原位根系(Stein et al., 2020) | Archaeopteris/Callixylon化石基干直径至少1.5 m,延伸至少10 m,重建的树高20~30 m,具大型树冠,次生木质部发达(Taylor et al., 2009) |
| 灌丛 | 早期真叶植物灌丛 | 加拿大魁北克下泥盆统Battery Point组,具木本真叶植物Kenrickia、Franhueberia等(Toledo et al., 2021) | 矿化保存的茎干片段,整体植株不明; Kenrickia茎干宽约7 mm(Toledo et al., 2021) |
| 湿地 | 早期似真蕨类湿地 | 美国东部上泥盆统Hampshire组,成煤植物主体为Rhacophyton(Scheckler,1986) | 重建的Rhacophyton植株高约1 m,具次生木质部,具大型侧枝或羽片系统(大于50 cm)(Taylor et al., 2009) |
), ArticleFig(id=1309812611047325934, tenantId=1146029695717560320, journalId=1308397449582313577, articleId=1309812603862483095, language=CN, label=表 1, caption=
志留纪—泥盆纪若干典型植被类型*
, figureFileSmall=null, figureFileBig=null, tableContent=
| 植被型组 | 植被类型 | 产地、层位及植被组成 | 可能的生物量信息 |
| 草地 | 莱尼蕨类草地 | 英国Anglo-Welsh盆地罗德洛统Pterinea层,具多种Cooksonia(Edwards and Richardson,2004) | Gerrienne等(2006)发现1枚近完整保存的Cooksonia标本,显示其植株高33 mm,茎轴宽度0.6~1.0 mm,多次分枝 |
| 莱尼蕨类草地 | 英国Anglo-Welsh盆地下泥盆统Ditton组,具多种Cooksonia、Salopella、Tortilicaulis等(Edwards and Richardson,2004) | 如上 |
| 工蕨类草地 | 加拿大极区Bathurst岛罗德洛统,具cf. Bathurstia sp.、Zosterophyllum sp.等(Kotyk et al., 2002) | Hao等(2010)发现1枚近完整保存的Zosterophyllum标本,显示植株地上部分高98 mm,根系统约20 mm,生物量根茎比约为0.028 |
| 工蕨类草地 | 贵州都匀下泥盆统蟒山群,具Zosterophyllum australianum、Z. baoyangense,混生Adoketophyton、Hedeia等(Huang et al., 2025) | 如上 |
| 工蕨类—早期真叶植物
混生草地 | 云南文山下泥盆统坡松冲组,工蕨纲物种占比43%,早期真叶植物物种占比22%,混生其他种类(Hao and Xue,2013) | 据估算,工蕨类Discalis植株高约20~30 cm,始叶蕨Eophyllophyton植株高约50~80 cm(Hao and Beck,1993;Hao and Xue,2013) |
| 工蕨类—镰蕨类
混生草地 | 云南曲靖下泥盆统徐家冲组,Zosterophyllum和Drepanophycus极为常见,同时混生Hedeia、Huia等(Wang et al., 2002;Xue et al., 2016) | 据Schweitzer(1980),Drepanophycus植株地上部分高50~60 cm;据Xue等(2016),Drepanophycus具复杂、庞大的地下茎系统 |
| 原始鳞木类草地 | 云南曲靖中泥盆统海口组,一些层位具密集、近原位的原始鳞木类植物Minarodendron(郑高峰等,2026) | 郑高峰等(2026)报道Minarodendron茎干最长可达1 m,宽4~5 mm,可能匍匐生长 |
| 森林 | 枝蕨类森林 | 美国纽约州中泥盆统Moscow组,森林主体为Eospermatopteris树木,兼具无脉树目、石松类分子(Stein et al., 2012) | Eospermatopteris树木基部宽50~150 cm,高至少8 m,侧枝逐渐脱落,仅树冠留存;在1200 m2的原位森林样方中,约具200个Eospermatopteris树以及其他植物种类(Stein et al., 2012) |
| 石松类森林 | 安徽广德上泥盆统五通组,森林主体为Guangdedendron树木(“新杭森林”),兼具早期似真蕨类等(Wang et al., 2019;Yang et al., 2024) | “新杭森林”为泥盆纪出露面积最大(至少250 000 m2)的森林,主体为Guangdedendron树木,估计高1.1~7.7 m,但大多低于3.2 m高,密度达38株/m2(Wang et al., 2019) |
| 前裸子植物森林 | Archaeopteris/Callixylon在晚泥盆世全球广布;美国纽约州中泥盆统Plattekill组,发现较为罕见的Archaeopteris原位根系(Stein et al., 2020) | Archaeopteris/Callixylon化石基干直径至少1.5 m,延伸至少10 m,重建的树高20~30 m,具大型树冠,次生木质部发达(Taylor et al., 2009) |
| 灌丛 | 早期真叶植物灌丛 | 加拿大魁北克下泥盆统Battery Point组,具木本真叶植物Kenrickia、Franhueberia等(Toledo et al., 2021) | 矿化保存的茎干片段,整体植株不明; Kenrickia茎干宽约7 mm(Toledo et al., 2021) |
| 湿地 | 早期似真蕨类湿地 | 美国东部上泥盆统Hampshire组,成煤植物主体为Rhacophyton(Scheckler,1986) | 重建的Rhacophyton植株高约1 m,具次生木质部,具大型侧枝或羽片系统(大于50 cm)(Taylor et al., 2009) |
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