Article(id=1309575093232165344, tenantId=1146029695717560320, journalId=1308397196074385509, issueId=1309575072856240766, articleNumber=null, orderNo=null, doi=10.11733/j.issn.1007-0435.2026.06.012, pmid=null, cstr=null, oa=null, hot=0, price=null, onlineType=0, articleFormat=0, articleType=null, articleTypeStr=null, receivedDate=1760371200000, receivedDateStr=2025-10-14, revisedDate=1767369600000, revisedDateStr=2026-01-03, acceptedDate=null, acceptedDateStr=null, onlineDate=1790156632954, onlineDateStr=2026-09-23, pubDate=null, pubDateStr=null, doiRegisterDate=null, doiRegisterDateStr=null, onlineIssueDate=1790156632954, onlineIssueDateStr=2026-09-23, onlineJustAcceptDate=null, onlineJustAcceptDateStr=null, onlineFirstDate=null, onlineFirstDateStr=null, sourceXml=null, magXml=null, createTime=1790156632954, creator=13701087609, updateTime=1790156632954, updator=13701087609, issue=Issue{id=1309575072856240766, tenantId=1146029695717560320, journalId=1308397196074385509, year='2026', volume='34', issue='6', pageStart='1963', pageEnd='2340', issueExtLink='null', onlineDate='null', pubDate='1781452800000', pubDateStr='2026-06-15', beforeIssueId=null, nextIssueId=null, price=null, status=1, issueComplete=1, articleOrder=1, issueType=-1, specialIssue=null, createTime=1790156628097, creator='13701087609', updateTime=1790156954064, updator='13701087609', preIssue=null, nextIssue=null, articleTotal=null, ext={EN=IssueExt(id=1309576440438419817, tenantId=1146029695717560320, journalId=1308397196074385509, issueId=1309575072856240766, language=EN, specialIssueTitle=, coverIllustrator=null, specialIssueEditor=, specialIssueAbout=), CN=IssueExt(id=1309576440438419818, tenantId=1146029695717560320, journalId=1308397196074385509, issueId=1309575072856240766, language=CN, specialIssueTitle=, coverIllustrator=null, specialIssueEditor=, specialIssueAbout=)}, issueFiles=null, downloadFileDto=null}, startPage=2090, endPage=2098, ext={EN=ArticleExt(id=1309575094964412899, articleId=1309575093232165344, tenantId=1146029695717560320, journalId=1308397196074385509, language=EN, title=Research on the Nitrogen and Phosphorus Homeostasis of Plant Community of Desert Steppe under Different Stocking Rates, columnId=null, journalTitle=Acta Agrestia Sinica, columnName=null, runingTitle=null, highlight=null, articleAbstract=In order to study the nitrogen and phosphorus homoeostasis of plant community of desert steppe under different stocking rates, the Stipa breviflora desert steppe in Inner Mongolia was used as the research object. The grazing experiment adopted a completely randomized block design, with a total of 4 levels of stocking rate and 3 repetitions, namely control (CK), light grazing (LG), moderate grazing (MG) and heavy grazing (HG) [sheep units·hm-2·0.5a-1]. The total nitrogen (TN) and the total phosphorus (TP) in plant community and soil were measured, and than calculated the homeostasis index (H), the results showed that the interaction between grazing and soil layer had no significant effect on the soil. Grazing attenuated TN and TP homeostatic regulation of the plant community. When the stocking rate was <1.1 or> 3.65, TN homeostasis was weakly homeostatic. The TP homeostasis was enhanced in HG. When the stocking rate was <1.09 or> 2.78, TP homeostasis was weakly homeostatic. The N∶P homeostasis was the strongest in MG, and when the stocking rate ranged from 0.17 to 1.9, the N∶P homeostasis was weakly homeostatic. Overall, when the stocking rate ranged from 0.17 to 1.09, which is a light to moderate grazing level, the grassland ecosystem had homeostatic regulation. The stocking rate has a minimum threshold of 2.37 for HTN and 1.94 for HTP, while a maximum threshold of 1.03 exists for HN∶P., authors=WANG Rong1, LÜ Shi-jie1, ZHANG Bo-han1, QU Zhi-qiang1, LI Zhi-guo1, authorsList=WANG Rong, LÜ Shi-jie, ZHANG Bo-han, QU Zhi-qiang, LI Zhi-guo, authorCompany=1 College of Grass Industry, Inner Mongolia Agricultural University/Key Laboratory of Grassland Resources, Ministry of Education, Hohhot, Inner Mongolia 010011, China, correspAuthors=null, authorNote=null, correspAuthorsNote=null, copyrightStatement=null, copyrightOwner=null, extLink=null, articleAbsUrl=null, sourceXml=null, magXml=null, pdfUrl=null, pdf=null, pdfFileSize=null, pdfExtLink=null, richHtmlUrl=null, mobilePdfUrl=null, reviewReport=null, pdfFirstPage=null, abstractGraph=null, abstractGraphContent=null, abstractVideo=null, citation=null, cebUrl=null, magXmlContent=null, mapNumber=null, fund=null), CN=ArticleExt(id=1309575094565954018, articleId=1309575093232165344, tenantId=1146029695717560320, journalId=1308397196074385509, language=CN, title=不同载畜率下荒漠草原植物群落氮磷内稳性研究, columnId=1309565024058835950, journalTitle=草地学报, columnName=研究论文, runingTitle=null, highlight=null, articleAbstract=为研究不同载畜率下荒漠草原植物群落氮磷内稳性特征,本文以内蒙古荒漠草原为研究对象,采取完全随机区组设计,共设置4个载畜率水平,3次重复,分别为对照组(Control,CK)0、轻度(Light stocking rate,LG)0.19、中度(Moderate stocking rate,MG)1.82和重度(Heavy stocking rate,HG)2.71[羊单位·hm-2·(0.5a)-1]。通过测定植物群落和土壤全氮(Total nitrogen,TN)、全磷(Total phosphorus,TP)含量,并计算内稳性指数(Homeostasis index,H),结果显示:放牧与土层的交互作用对土壤无显著影响。放牧降低了植物群落TN和TP含量及其内稳性,当载畜率<1.1或>3.65时,TN内稳性为弱稳态。TP内稳性在HG区有所增强,当载畜率<1.09或>2.78时TP为弱稳态。MG区N∶P内稳性最强,当载畜率为0.17~1.9时N∶P内稳性为弱稳态。总体来看,当载畜率为0.17~1.09,为轻中度放牧时,草地生态系统均具有一定的稳态调节能力。载畜率对HTN和HTP均存在一个最小阈值2.37和1.94;对HN∶P存在一个最大阈值1.03。, authors=汪蓉1, 吕世杰1, 张博涵1, 屈志强1, 李治国1, authorsList=汪蓉, 吕世杰, 张博涵, 屈志强, 李治国, authorCompany=1 内蒙古农业大学草业学院/草地资源教育部重点实验室, 内蒙古 呼和浩特 010011, correspAuthors=null, authorNote=汪蓉(1999-),女,汉族,青海海东人,硕士研究生,主要从事草地资源与管理研究,E-mail:wr19490@163.com, correspAuthorsNote=null, copyrightStatement=null, copyrightOwner=null, extLink=null, articleAbsUrl=null, sourceXml=null, magXml=null, pdfUrl=null, pdf=yiXSOKnYNKkPdQ0VFXERbQ==, pdfFileSize=1987121, pdfExtLink=null, richHtmlUrl=null, mobilePdfUrl=null, reviewReport=null, pdfFirstPage=null, abstractGraph=null, abstractGraphContent=null, abstractVideo=null, citation=null, cebUrl=null, magXmlContent=null, mapNumber=null, fund=国家重点研发项目-基于生态和经济双赢的草地-家畜系统可持续管理研究 (2024YFE0113100); 内蒙古自治区“揭榜挂帅”项目“荒漠草原综合治理与家庭牧场可持续管理集成技术示范” (2024JBGS0011); 内蒙古自治区一流学科科研专项项目 (YLXKZX-NND-029); 国家自然科学基金项目 (32060384))}, authors=[Author(id=1320569737063547782, tenantId=1146029695717560320, journalId=null, articleId=1309575093232165344, orderNo=null, firstName=null, middleName=null, lastName=null, nameCn=null, orcid=null, stid=null, country=null, authorPic=null, dead=null, email=null, emailSecond=null, emailThird=null, correspondingAuthor=null, authorType=null, ext={EN=AuthorExt(id=null, tenantId=null, 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Ecological stoichiometry: from sea to lake to land[J]. Trends in Ecology & Evolution,2000,15(10):393-394 KOOJIMAN S A L M. The stoichiometry of animal energetics[J]. Journal of Theoretical Biology,1995,177(2):139-149 ZENG D H, CHENG G S. Ecological stoichiometry: a science to explore the complexity of living systems[J]. Acta Phytoecologica Sinica,2005,29(6):1007-1019 曾德慧,陈广生. 生态化学计量学:复杂生命系统奥秘的探索[J]. 植物生态学报,2005,29(6):1007-1019 JEYASINGH P D,WEIDER L J,STERNER R W. Genetically-based trade-offs in response to stoichiometric food quality influence competition in a keystone aquatic herbivore[J]. Ecology Letters,2009,12(11):1229-1237 CHEN Z F,ZHOU J J,LAI S B,et al. Species differences in stoichiometric homeostasis affect grassland community stability under N and P addition[J]. Environmental Science and Pollution Research,2023,30(22):61913-61926 HESSEN D O,ÅGREN G I,ANDERSON T R,et al. Carbon sequestration in ecosystems: the role of stoichiometry[J]. Ecology,2004,85(5):1179-1192 YU Q,CHEN Q S,ELSER J J,et al. Linking stoichiometric homoeostasis with ecosystem structure,functioning and stability[J]. Ecology Letters,2010,13(11):1390-1399 YU Q,ELSER J J,HE N P,et al. Stoichiometric homeostasis of vascular plants in the Inner Mongolia grassland[J]. Oecologia,2011,166(1):1-10 GÜSEWELL S. N: P ratios in terrestrial plants: variation and functional significance[J]. The New Phytologist,2004,164(2):243-266 KOERSELMAN W,MEULEMAN A F M. The vegetation N:P ratio: a new tool to detect the nature of nutrient limitation[J]. The Journal of Applied Ecology,1996,33(6):1441 WANG M,ZHANG Y,LI R Q,et al. Effects of grazing disturbance on the stoichiometry of nitrogen and phosphorus in plant organs of Leymus chinensis meadow steppe[J]. Scientia Agricultura Sinica,2022,55(7):1371-1384 王淼,张宇,李瑞强,等. 放牧强度对羊草草甸草原植物器官及群落氮磷化学计量的影响[J], 中国农业科学,2022,55(7):1371-1384 CHEN H J. The response of reproductive characteristics and ecological stoichiometry of main plant population to stocking rate in Inner Mongolia desert steppe[D]. Hohhot:Inner Mongolia Agricultural University,2011:116-119 陈海军. 荒漠草原主要植物种群繁殖性状及化学计量特征对载畜率的响应[D]. 呼和浩特:内蒙古农业大学,2011:116-119 STERNER R W,ELSER J J. Ecological Stoichiometry: The Biology of Elements From Molecules to The Biosphere[M]. Princeton, New Jersey:Princeton University Press,2002:15-18 PERSSON J,FINK P,GOTO A,et al. To be or not to be what you eat: regulation of stoichiometric homeostasis among autotrophs and heterotrophs[J]. Oikos,2010,119(5):741-751 YANG L,MA X Z,LI Y Q. Effects of grazing on ecological stoichiometry of Stipa krylovii and soil in desert grassland[J]. Acta Botanica Boreali-Occidentalia Sinica,2020,40(2):328-334 杨林,马秀枝,李依倩. 放牧对荒漠草原克氏针茅种群和土壤生态化学计量特征的影响[J], 西北植物学报,2020,40(2):328-334 HASSAN N,LI X F,WANG J Y,et al. Effects of grazing on C∶N∶P stoichiometry attenuate from soils to plants and insect herbivores in a semi-arid grassland[J]. Oecologia,2021,195(3):785-795 ZHOU Q,JIANG Y B,HAO J H,et al. Advances in the study of biogeochemical cycles of phosphorus[J]. Geological Journal of China Universities,2021,27(2):183-199 周强,姜允斌,郝记华,等. 磷的生物地球化学循环研究进展[J]. 高校地质学报,2021,27(2):183-199 LI Y Q. Effect of different stocking rates on the characteristic and mechanism of soil wind erosion in a Stipa breviflora desert steppe[D]. Hohhot:Inner Mongolia Agricultural University,2016:8-10 李永强. 不同载畜率对短花针茅荒漠草原土壤风蚀的影响及作用机制[D]. 呼和浩特:内蒙古农业大学,2016:8-10 ZHAO M M. Spatial distribution of soil physical-chemical characteristics and erodibility of desert steppe with different grazing intensities[D]. Taian:Shandong Agricultural University,2022:40-41 赵美曼. 荒漠草原不同放牧强度土壤理化特性及可蚀性的空间分布[D]. 泰安:山东农业大学,2022:40-41 AN Y,AN H,LI S B. Effects of grazing on ecological stoichiometry of soil and dominant plants in desert grassland[J]. Acta Prataculturae Sinica,2018,27(12):94-102 安钰,安慧,李生兵. 放牧对荒漠草原土壤和优势植物生态化学计量特征的影响[J]. 草业学报,2018,27(12):94-102 DING X H,GONG L,WANG D B,et al. Grazing effects on eco-stoichiometry of plant and soil in Hulunbeir,Inner Mogolia[J]. Acta Ecologica Sinica,2012,32(15):4722-4730 丁小慧,宫立,王东波,等. 放牧对呼伦贝尔草地植物和土壤生态化学计量学特征的影响[J]. 生态学报,2012,32(15):4722-4730 LU X X,ZHANG W Q,WANG H R,et al. Effects of grazing intensity on soil phosphorus in typical grassland of Inner Mongolia[J]. Journal of Arid Land Resources and Environment,2022,36(5):153-159 卢晓霞,张卫青,王海茹,等. 放牧强度对内蒙古典型草原土壤磷素的影响[J]. 干旱区资源与环境,2022,36(5):153-159 LI L. Effects of Tan-sheep grazing on C: N: P stoichiometry and multifunctionality of typical steppe[D]. Lanzhou:Lanzhou University,2021:91-96 李岚. 滩羊放牧对典型草原生态化学计量特征和多功能性的影响[D]. 兰州:兰州大学,2021:91-96 GÜSEWELL S. Variation in nitrogen and phosphorus concentrations of wetland plants[J]. Perspectives in Plant Ecology,Evolution and Systematics,2002,5(1):37-61 DING L J,WEI Z J,ZHANG S,et al. Effects of seasonal regulation of grazing intensity on Soil and plant nutrients in desert steppe[J]. Grassland and Prataculture,2019,31(1):53-59 丁莉君,卫智军,张爽,等. 放牧强度季节调控对荒漠草原土壤和植物养分的影响[J]. 草原与草业,2019,31(1):53-59 SUN C. Study on grassland degeneration based on ecological stoichiometry[D]. Changchun:Jilin University,2012:39-42 孙超. 基于生态化学计量学的草地退化研究[D]. 长春:吉林大学,2012:39-42 GAO Q J,ZHU W Y,HOU J J,et al. Effects of grazing intensity on ecostoichiometric characteristics of plant leaves in Alpine meadow[J]. Chinese Journal of Grassland,2019,41(3):45-50 高巧静,朱文琰,侯将将,等. 放牧强度对高寒草甸植物叶片生态化学计量特征的影响[J]. 中国草地学报,2019,41(3):45-50 WANG Y L,ZHU W Y,HOU J J,et al. Effects of grazing on leaf traits of four degraded indicator plants in Guinan[J]. Pratacultural Science,2020,37(3):423-431 王娅琳,朱文琰,侯将将,等. 放牧对贵南县4种退化指示植物叶片性状的影响[J]. 草业科学,2020,37(3):423-431 HE M,XU Q Y,XIA Y,et al. Plant phosphorus acquisition mechanisms and their response to global climate changes[J]. Chinese Journal of Plant Ecology,2023,47(3):291-305 何敏,许秋月,夏允,等. 植物磷获取机制及其对全球变化的响应[J]. 植物生态学报,2023,47(3):291-305 YU Q,WILCOX K,LA PIERRE K,et al. Stoichiometric homeostasis predicts plant species dominance,temporal stability,and responses to global change[J]. Ecology,2015,96(9):2328-2335 WANG X T,CHAI J,JIANG C,et al. Population spatial pattern of Stipa grandis and its response to long-term overgrazing[J]. Biodiversity Science,2020,28(2):128-134 王鑫厅,柴静,姜超,等. 典型草原大针茅种群空间格局及对长期过度放牧的响应[J]. 生物多样性,2020,28(2):128-134 ZHANG N,QIN Y,JIN K,et al. Effects of grazing on vegetation community and soil physical properties in typical steppe[J]. Chinese Journal of Grassland,2020,42(4):91-100 张娜,秦艳,金轲,等. 放牧对典型草原群落特征及土壤物理性状的影响[J]. 中国草地学报,2020,42(4):91-100 CHENG Y Y. Study on the relationship between stoichiometric homeostasis and stability of forest ecosystem invaded by moso bamboo[D]. Hangzhou:Zhejiang A & F University,2014:4-10 程艳艳. 化学计量内稳性与毛竹入侵生态系统稳定性关系的研究[D]. 杭州:浙江农林大学,2014:4-10 HOOVER D L,KNAPP A K,SMITH M D. Resistance and resilience of a grassland ecosystem to climate extremes[J]. Ecology,2014,95(9):2646-2656 HU Y F,LIU J Y,ZHUANG D F. Progress in wind erosion research[J]. Progress in Geography,2003,22(3):288-295 胡云锋,刘纪远,庄大方. 土壤风力侵蚀研究现状与进展[J]. 地理科学进展,2003,22(3):288-295)
Research on the Nitrogen and Phosphorus Homeostasis of Plant Community of Desert Steppe under Different Stocking Rates
WANG Rong, LÜ Shi-jie, ZHANG Bo-han, QU Zhi-qiang, LI Zhi-guo
Affiliations
1 College of Grass Industry, Inner Mongolia Agricultural University/Key Laboratory of Grassland Resources, Ministry of Education, Hohhot, Inner Mongolia 010011, China
In order to study the nitrogen and phosphorus homoeostasis of plant community of desert steppe under different stocking rates, the Stipa breviflora desert steppe in Inner Mongolia was used as the research object. The grazing experiment adopted a completely randomized block design, with a total of 4 levels of stocking rate and 3 repetitions, namely control (CK), light grazing (LG), moderate grazing (MG) and heavy grazing (HG) [sheep units·hm-2·0.5a-1]. The total nitrogen (TN) and the total phosphorus (TP) in plant community and soil were measured, and than calculated the homeostasis index (H), the results showed that the interaction between grazing and soil layer had no significant effect on the soil. Grazing attenuated TN and TP homeostatic regulation of the plant community. When the stocking rate was <1.1 or> 3.65, TN homeostasis was weakly homeostatic. The TP homeostasis was enhanced in HG. When the stocking rate was <1.09 or> 2.78, TP homeostasis was weakly homeostatic. The N∶P homeostasis was the strongest in MG, and when the stocking rate ranged from 0.17 to 1.9, the N∶P homeostasis was weakly homeostatic. Overall, when the stocking rate ranged from 0.17 to 1.09, which is a light to moderate grazing level, the grassland ecosystem had homeostatic regulation. The stocking rate has a minimum threshold of 2.37 for HTN and 1.94 for HTP, while a maximum threshold of 1.03 exists for HN∶P.
WANG Rong, LÜ Shi-jie, ZHANG Bo-han, QU Zhi-qiang, LI Zhi-guo.
Research on the Nitrogen and Phosphorus Homeostasis of Plant Community of Desert Steppe under Different Stocking Rates[J].
Acta Agrestia Sinica,
2026
, 34
(6)
: 2090
-2098
.
DOI: 10.11733/j.issn.1007-0435.2026.06.012
ELSER J J. Ecological stoichiometry: from sea to lake to land[J]. Trends in Ecology & Evolution,2000,15(10):393-394 KOOJIMAN S A L M. The stoichiometry of animal energetics[J]. Journal of Theoretical Biology,1995,177(2):139-149 ZENG D H, CHENG G S. Ecological stoichiometry: a science to explore the complexity of living systems[J]. Acta Phytoecologica Sinica,2005,29(6):1007-1019 曾德慧,陈广生. 生态化学计量学:复杂生命系统奥秘的探索[J]. 植物生态学报,2005,29(6):1007-1019 JEYASINGH P D,WEIDER L J,STERNER R W. Genetically-based trade-offs in response to stoichiometric food quality influence competition in a keystone aquatic herbivore[J]. Ecology Letters,2009,12(11):1229-1237 CHEN Z F,ZHOU J J,LAI S B,et al. Species differences in stoichiometric homeostasis affect grassland community stability under N and P addition[J]. Environmental Science and Pollution Research,2023,30(22):61913-61926 HESSEN D O,ÅGREN G I,ANDERSON T R,et al. Carbon sequestration in ecosystems: the role of stoichiometry[J]. Ecology,2004,85(5):1179-1192 YU Q,CHEN Q S,ELSER J J,et al. Linking stoichiometric homoeostasis with ecosystem structure,functioning and stability[J]. Ecology Letters,2010,13(11):1390-1399 YU Q,ELSER J J,HE N P,et al. Stoichiometric homeostasis of vascular plants in the Inner Mongolia grassland[J]. Oecologia,2011,166(1):1-10 GÜSEWELL S. N: P ratios in terrestrial plants: variation and functional significance[J]. The New Phytologist,2004,164(2):243-266 KOERSELMAN W,MEULEMAN A F M. The vegetation N:P ratio: a new tool to detect the nature of nutrient limitation[J]. The Journal of Applied Ecology,1996,33(6):1441 WANG M,ZHANG Y,LI R Q,et al. Effects of grazing disturbance on the stoichiometry of nitrogen and phosphorus in plant organs of Leymus chinensis meadow steppe[J]. Scientia Agricultura Sinica,2022,55(7):1371-1384 王淼,张宇,李瑞强,等. 放牧强度对羊草草甸草原植物器官及群落氮磷化学计量的影响[J], 中国农业科学,2022,55(7):1371-1384 CHEN H J. The response of reproductive characteristics and ecological stoichiometry of main plant population to stocking rate in Inner Mongolia desert steppe[D]. Hohhot:Inner Mongolia Agricultural University,2011:116-119 陈海军. 荒漠草原主要植物种群繁殖性状及化学计量特征对载畜率的响应[D]. 呼和浩特:内蒙古农业大学,2011:116-119 STERNER R W,ELSER J J. Ecological Stoichiometry: The Biology of Elements From Molecules to The Biosphere[M]. Princeton, New Jersey:Princeton University Press,2002:15-18 PERSSON J,FINK P,GOTO A,et al. To be or not to be what you eat: regulation of stoichiometric homeostasis among autotrophs and heterotrophs[J]. Oikos,2010,119(5):741-751 YANG L,MA X Z,LI Y Q. Effects of grazing on ecological stoichiometry of Stipa krylovii and soil in desert grassland[J]. Acta Botanica Boreali-Occidentalia Sinica,2020,40(2):328-334 杨林,马秀枝,李依倩. 放牧对荒漠草原克氏针茅种群和土壤生态化学计量特征的影响[J], 西北植物学报,2020,40(2):328-334 HASSAN N,LI X F,WANG J Y,et al. Effects of grazing on C∶N∶P stoichiometry attenuate from soils to plants and insect herbivores in a semi-arid grassland[J]. Oecologia,2021,195(3):785-795 ZHOU Q,JIANG Y B,HAO J H,et al. Advances in the study of biogeochemical cycles of phosphorus[J]. Geological Journal of China Universities,2021,27(2):183-199 周强,姜允斌,郝记华,等. 磷的生物地球化学循环研究进展[J]. 高校地质学报,2021,27(2):183-199 LI Y Q. Effect of different stocking rates on the characteristic and mechanism of soil wind erosion in a Stipa breviflora desert steppe[D]. Hohhot:Inner Mongolia Agricultural University,2016:8-10 李永强. 不同载畜率对短花针茅荒漠草原土壤风蚀的影响及作用机制[D]. 呼和浩特:内蒙古农业大学,2016:8-10 ZHAO M M. Spatial distribution of soil physical-chemical characteristics and erodibility of desert steppe with different grazing intensities[D]. Taian:Shandong Agricultural University,2022:40-41 赵美曼. 荒漠草原不同放牧强度土壤理化特性及可蚀性的空间分布[D]. 泰安:山东农业大学,2022:40-41 AN Y,AN H,LI S B. Effects of grazing on ecological stoichiometry of soil and dominant plants in desert grassland[J]. Acta Prataculturae Sinica,2018,27(12):94-102 安钰,安慧,李生兵. 放牧对荒漠草原土壤和优势植物生态化学计量特征的影响[J]. 草业学报,2018,27(12):94-102 DING X H,GONG L,WANG D B,et al. Grazing effects on eco-stoichiometry of plant and soil in Hulunbeir,Inner Mogolia[J]. Acta Ecologica Sinica,2012,32(15):4722-4730 丁小慧,宫立,王东波,等. 放牧对呼伦贝尔草地植物和土壤生态化学计量学特征的影响[J]. 生态学报,2012,32(15):4722-4730 LU X X,ZHANG W Q,WANG H R,et al. Effects of grazing intensity on soil phosphorus in typical grassland of Inner Mongolia[J]. Journal of Arid Land Resources and Environment,2022,36(5):153-159 卢晓霞,张卫青,王海茹,等. 放牧强度对内蒙古典型草原土壤磷素的影响[J]. 干旱区资源与环境,2022,36(5):153-159 LI L. Effects of Tan-sheep grazing on C: N: P stoichiometry and multifunctionality of typical steppe[D]. Lanzhou:Lanzhou University,2021:91-96 李岚. 滩羊放牧对典型草原生态化学计量特征和多功能性的影响[D]. 兰州:兰州大学,2021:91-96 GÜSEWELL S. Variation in nitrogen and phosphorus concentrations of wetland plants[J]. Perspectives in Plant Ecology,Evolution and Systematics,2002,5(1):37-61 DING L J,WEI Z J,ZHANG S,et al. Effects of seasonal regulation of grazing intensity on Soil and plant nutrients in desert steppe[J]. Grassland and Prataculture,2019,31(1):53-59 丁莉君,卫智军,张爽,等. 放牧强度季节调控对荒漠草原土壤和植物养分的影响[J]. 草原与草业,2019,31(1):53-59 SUN C. Study on grassland degeneration based on ecological stoichiometry[D]. Changchun:Jilin University,2012:39-42 孙超. 基于生态化学计量学的草地退化研究[D]. 长春:吉林大学,2012:39-42 GAO Q J,ZHU W Y,HOU J J,et al. Effects of grazing intensity on ecostoichiometric characteristics of plant leaves in Alpine meadow[J]. Chinese Journal of Grassland,2019,41(3):45-50 高巧静,朱文琰,侯将将,等. 放牧强度对高寒草甸植物叶片生态化学计量特征的影响[J]. 中国草地学报,2019,41(3):45-50 WANG Y L,ZHU W Y,HOU J J,et al. Effects of grazing on leaf traits of four degraded indicator plants in Guinan[J]. Pratacultural Science,2020,37(3):423-431 王娅琳,朱文琰,侯将将,等. 放牧对贵南县4种退化指示植物叶片性状的影响[J]. 草业科学,2020,37(3):423-431 HE M,XU Q Y,XIA Y,et al. Plant phosphorus acquisition mechanisms and their response to global climate changes[J]. Chinese Journal of Plant Ecology,2023,47(3):291-305 何敏,许秋月,夏允,等. 植物磷获取机制及其对全球变化的响应[J]. 植物生态学报,2023,47(3):291-305 YU Q,WILCOX K,LA PIERRE K,et al. Stoichiometric homeostasis predicts plant species dominance,temporal stability,and responses to global change[J]. Ecology,2015,96(9):2328-2335 WANG X T,CHAI J,JIANG C,et al. Population spatial pattern of Stipa grandis and its response to long-term overgrazing[J]. Biodiversity Science,2020,28(2):128-134 王鑫厅,柴静,姜超,等. 典型草原大针茅种群空间格局及对长期过度放牧的响应[J]. 生物多样性,2020,28(2):128-134 ZHANG N,QIN Y,JIN K,et al. Effects of grazing on vegetation community and soil physical properties in typical steppe[J]. Chinese Journal of Grassland,2020,42(4):91-100 张娜,秦艳,金轲,等. 放牧对典型草原群落特征及土壤物理性状的影响[J]. 中国草地学报,2020,42(4):91-100 CHENG Y Y. Study on the relationship between stoichiometric homeostasis and stability of forest ecosystem invaded by moso bamboo[D]. Hangzhou:Zhejiang A & F University,2014:4-10 程艳艳. 化学计量内稳性与毛竹入侵生态系统稳定性关系的研究[D]. 杭州:浙江农林大学,2014:4-10 HOOVER D L,KNAPP A K,SMITH M D. Resistance and resilience of a grassland ecosystem to climate extremes[J]. Ecology,2014,95(9):2646-2656 HU Y F,LIU J Y,ZHUANG D F. Progress in wind erosion research[J]. Progress in Geography,2003,22(3):288-295 胡云锋,刘纪远,庄大方. 土壤风力侵蚀研究现状与进展[J]. 地理科学进展,2003,22(3):288-295