Article(id=1309574771776520228, tenantId=1146029695717560320, journalId=1308397196074385509, issueId=1309574765791244449, articleNumber=null, orderNo=null, doi=10.11733/j.issn.1007-0435.2026.07.025, pmid=null, cstr=null, oa=null, hot=0, price=null, onlineType=0, articleFormat=0, articleType=null, articleTypeStr=null, receivedDate=1775664000000, receivedDateStr=2026-04-09, revisedDate=1778428800000, revisedDateStr=2026-05-11, acceptedDate=null, acceptedDateStr=null, onlineDate=1790156556313, onlineDateStr=2026-09-23, pubDate=null, pubDateStr=null, doiRegisterDate=null, doiRegisterDateStr=null, onlineIssueDate=1790156556313, onlineIssueDateStr=2026-09-23, onlineJustAcceptDate=null, onlineJustAcceptDateStr=null, onlineFirstDate=null, onlineFirstDateStr=null, sourceXml=null, magXml=null, createTime=1790156556313, creator=13701087609, updateTime=1790156556313, updator=13701087609, issue=Issue{id=1309574765791244449, tenantId=1146029695717560320, journalId=1308397196074385509, year='2026', volume='34', issue='7', pageStart='2341', pageEnd='2752', issueExtLink='null', onlineDate='null', pubDate='1784044800000', pubDateStr='2026-07-15', beforeIssueId=null, nextIssueId=null, price=null, status=1, issueComplete=1, articleOrder=1, issueType=-1, specialIssue=null, createTime=1790156554887, creator='13701087609', updateTime=1790156849638, updator='13701087609', preIssue=null, nextIssue=null, articleTotal=null, ext={EN=IssueExt(id=1309576002150424823, tenantId=1146029695717560320, journalId=1308397196074385509, issueId=1309574765791244449, language=EN, specialIssueTitle=, coverIllustrator=null, specialIssueEditor=, specialIssueAbout=), CN=IssueExt(id=1309576002154619128, tenantId=1146029695717560320, journalId=1308397196074385509, issueId=1309574765791244449, language=CN, specialIssueTitle=, coverIllustrator=null, specialIssueEditor=, specialIssueAbout=)}, issueFiles=null, downloadFileDto=null}, startPage=2624, endPage=2637, ext={EN=ArticleExt(id=1309574774012084262, articleId=1309574771776520228, tenantId=1146029695717560320, journalId=1308397196074385509, language=EN, title=Structural Regulation Mechanisms of Soil Aggregate Reorganization and Carbon Pool Responses to Grassland Degradation in Humid Regions, columnId=null, journalTitle=Acta Agrestia Sinica, columnName=null, runingTitle=null, highlight=null, articleAbstract=Grassland degradation is a major driver affecting soil structure stability and carbon cycling processes; however, its underlying pathways and regulatory mechanisms under humid climatic conditions remain insufficiently quantified. In this study, a typical montane meadow in Nanshan Pasture, Hunan Province, China, was selected to investigate the characteristics of soil aggregate structure and organic carbon distribution across a degradation gradient, including non-degraded (ND), lightly degraded (LD), moderately degraded (MD), and heavily degraded (HD) grasslands within the 0-30 cm soil layer. The results showed that, with increasing degradation intensity, the proportion of >5 mm macroaggregates decreased by approximately 47%, whereas the proportion of <0.25 mm microaggregates increased by approximately 168%. Meanwhile, the mean weight diameter (MWD) and geometric mean diameter (GMD) of both mechanically stable and water-stable aggregates declined by approximately 46% and 53%, respectively, accompanied by a significant increase in fractal dimension, indicating a transition of soil structure from a stable state to a fragmented state. The breakdown of macroaggregates released organic carbon, part of which was subsequently adsorbed and stabilized by microaggregates, resulting in a shift in carbon distribution from a “macroaggregate-dominated” pattern to a “microaggregate-dominated” pattern. Grassland degradation also led to a decline in total soil organic carbon and induced substantial structural reorganization of soil carbon pools, highlighting the pivotal role of soil aggregates in carbon protection and redistribution. These findings elucidate the structural mechanisms underlying carbon pool dynamics driven by grassland degradation in humid regions and provide a theoretical basis for ecological restoration and carbon sequestration management of degraded grasslands., authors=HUANG Hui-qun1, LIU Xi-yuan1, ZHU Xiao-hua1, ZENG He-ping2, authorsList=HUANG Hui-qun, LIU Xi-yuan, ZHU Xiao-hua, ZENG He-ping, authorCompany=1 Shaoyang Grassland Resources Protection Center, Shaoyang, Hunan Province 422000, China; 2 City college, Kunming University of Science and Technology, Kunming, Yunnan Province 650500, China, correspAuthors=null, authorNote=null, correspAuthorsNote=null, copyrightStatement=null, copyrightOwner=null, extLink=null, articleAbsUrl=null, sourceXml=null, magXml=null, pdfUrl=null, pdf=null, pdfFileSize=null, pdfExtLink=null, richHtmlUrl=null, mobilePdfUrl=null, reviewReport=null, pdfFirstPage=null, abstractGraph=null, abstractGraphContent=null, abstractVideo=null, citation=null, cebUrl=null, magXmlContent=null, mapNumber=null, fund=null), CN=ArticleExt(id=1309574773932392485, articleId=1309574771776520228, tenantId=1146029695717560320, journalId=1308397196074385509, language=CN, title=湿润区草地退化对土壤团聚体结构重组及碳库响应的结构调控机制, columnId=1309565024058835950, journalTitle=草地学报, columnName=研究论文, runingTitle=null, highlight=null, articleAbstract=草地退化是影响土壤结构稳定性与碳循环过程的重要驱动因素,但湿润气候条件下其作用路径及调控机制仍缺乏定量解析。本研究以湖南南山牧场典型山地草甸为对象,基于退化梯度(ND,LD,MD,HD),系统分析了0~30 cm土层土壤团聚体结构及有机碳分布特征。结果表明,随着退化程度加剧,>5 mm大团聚体含量下降约47%,<0.25 mm微团聚体含量增加约168%,机械稳定性和水稳性团聚体的平均重量直径(MWD)和几何平均直径(GMD)分别降低约46%,53%,分形维数显著升高,表明土壤结构由稳定态向破碎态转变。大团聚体破碎释放碳,其中一部分被微团聚体吸附封存,导致碳在团聚体中的分布由“大团聚体主导”向“微团聚体主导”转变。草地退化同时引起土壤有机碳总量下降与碳结构性重组,显示土壤团聚体在碳保护和分配中的关键作用。本研究揭示了湿润区草地退化驱动的碳库变化及其结构性机制,为退化草地生态修复与碳汇管理提供了理论依据。, authors=黄惠群1, 刘西苑1, 朱晓花1, 曾和平2, authorsList=黄惠群, 刘西苑, 朱晓花, 曾和平, authorCompany=1 邵阳市草地资源保护中心, 湖南 邵阳 422000; 2 昆明理工大学城市学院, 云南 昆明 650500, correspAuthors=null, authorNote=黄惠群(1989-),男,苗族,湖南邵阳人,硕士研究生,主要从事生态保护与修复、生态系统功能与生物多样性、植物-土壤-微生物互作、草种质资源收集与培育、林草技术推广等研究,E-mail:978344016@qq.com, correspAuthorsNote=null, copyrightStatement=null, copyrightOwner=null, extLink=null, articleAbsUrl=null, sourceXml=null, magXml=null, pdfUrl=null, pdf=sK+TK1qLoIiIf9qG+qSPCQ==, pdfFileSize=3807402, pdfExtLink=null, richHtmlUrl=null, mobilePdfUrl=null, reviewReport=null, pdfFirstPage=null, abstractGraph=null, abstractGraphContent=null, abstractVideo=null, citation=null, cebUrl=null, magXmlContent=null, mapNumber=null, fund=南山国家公园草地生态系统定位监测研究 (XLK202108-10); 国家自然科学基金项目 (41461061))}, authors=[Author(id=1320972827495354802, tenantId=1146029695717560320, journalId=null, articleId=1309574771776520228, orderNo=null, firstName=null, middleName=null, lastName=null, nameCn=null, orcid=null, stid=null, country=null, authorPic=null, dead=null, email=null, emailSecond=null, emailThird=null, correspondingAuthor=null, authorType=null, ext={EN=AuthorExt(id=null, tenantId=null, 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pdfUrlEn=https://castjournals.cast.org.cn/joweb/cdxb/EN/PDF/10.11733/j.issn.1007-0435.2026.07.025, aliStartDate=null, aliEndDate=null, collectionFlag=false, citedCount=null, citedUrl=null, previewStatus=0, delFlag=0, hasFullText=0, orderTime=1784044800000, fullTextJson=null, articleText=null, reference=YAHDJIAN L, CAMPANA S, TOGNETTI P M, et al. Insights on global rangeland ecosystem services shaped by grazing and fertilization[J]. Frontiers in Ecology and the Environment, 2026, 24(3): e70022 WAHEED A, QIAO X, MUHAMMAD M, et al. Plant root-mediated carbon sequestration and nutrient cycling in grassland ecosystems under land use and climate change[J]. Agriculture, Ecosystems & Environment, 2025, 393: 109865 GUO H N, XIN X P, CHEN J Q, et al. Divergent responses of particulate and mineral-associated organic carbon stock to grazing in a Eurasian temperate meadow steppe[J]. Geoderma, 2025, 462: 117516 LI H N, MENG R B, MENG Z J, et al. Influence of grazing patterns on the stability of soil aggregates in semi-arid grasslands[J]. Journal of Arid Land, 2026, 18(2): 322-338 VAN DAEL T, VERWERFT Q, SMOLDERS E. The soil structural stability determined by the QuantiSlakeTest: Linkage with soil porosity, water-stable aggregate fractions and soil chemical properties[J]. Geoderma, 2025, 457: 117281 HU P L, ZHANG W, NOTTINGHAM A T, et al. Lithological controls on soil aggregates and minerals regulate microbial carbon use efficiency and necromass stability[J]. Environmental Science & Technology, 2024, 58(48): 21186-21199 QIN W P, YUAN H, SUN M Y, et al. Effects of Grazing Exclusion on Soil Aggregate Composition and Stability in Degraded Alpine Meadows[J/OL]. https://doi.org/10.19741/j.issn.1673-4831.2025.0554, 2025-10-29/2026-04-09 秦文萍, 袁晖, 孙铭阳, 等. 围栏封育对退化高寒草甸土壤团聚体组成及稳定性的影响[J/OL]. https://doi.org/10.19741/j.issn.1673-4831.2025.0554, 2025-10-29/2026-04-09 DU Z Y, CONG N, ZHAO G, et al. Divergent responses of vegetation and soil characteristics to grassland degradation in the Qinghai-Tibet and Inner Mongolia plateaus[J]. Catena, 2025, 257: 109146 LIN X, ZHANG S W, HUETE A, et al. Global-to-regional variations in the effects of grassland management on soil carbon[J]. Agriculture, Ecosystems & Environment, 2026, 400: 110251 ZHOU L, SU Z A, YANG J, et al. Effects of soil erosion on aggregate -associated organic carbon in typical grassland and terraced fields of the agro -pastoral ecotone of Qilian Mountains[J]. Journal of Soil and Water Conservation, 2025, 39(4): 198-207 周铃, 苏正安, 杨静, 等. 祁连山农牧交错带典型草地和梯田土壤侵蚀对团聚体有机碳的影响[J]. 水土保持学报, 2025, 39(4): 198-207 MA Q, BAI W, MOU G X, et al. Effects of warming on soil Aggregate stability and contents of carbon and nitrogen in Alpine swamp meadow [J]. Acta Agrestia Sinica, 2025, 33(2): 547-555 马群, 白炜, 牟国旭, 等. 增温对高寒沼泽草甸土壤团聚体稳定性及碳氮含量的影响[J]. 草地学报, 2025, 33(2): 547-555 MI J, WANG F, SHI J, et al. Contrasting trends in plant diversity and soil carbon mineralization under precipitation-driven vegetation and soil carbon dynamics in the Mongolian Plateau[J]. Ecology and Evolution, 2025, 15(7): e71806 LUO J Y. Driving mechanism of biochar combined enclosure on multifunctional restoration of degraded grassland ecosystem in Nanshan pasture[D]. Changsha:Central South University of Forestry & Technology, 2025:11 罗俊毅. 生物炭联合围封对南山牧场退化草地生态系统多功能恢复的驱动机制[D]. 长沙:中南林业科技大学, 2025:11 ZHANG W. Nanshan Grassland, where Laoshanjie is located[N]. China Cultural Relics News, 2023-02-21(1) 张伟. 老山界所在地——南山草原[N]. 中国文物报, 2023-02-21(1) YOU L S, LIU S M, XIAO T G, et al. Investigation on plant and insect species in Nanshan pasture farm, Hunan Province and insect fauna analysis[J].Journal of Hunan Agricultural University (Natural Sciences), 1999, 25(1): 47-51 游兰韶, 刘寿明, 肖铁光, 等. 湖南省南山牧场植物和昆虫种类调查及区系分析[J]. 湖南农业大学学报, 1999, 25(1): 47-51 ZHANG Y. Study on the changes and influencing factors of soil C∶N∶P in different grassland types in Nanshan, Hunan Province[D]. Changsha:Central South University of Forestry & Technology, 2023:9 张宇. 湖南南山不同草地类型土壤C∶N∶P变化及其影响因素研究[D]. 长沙:中南林业科技大学, 2023:9 LI Y J. The grassland condition of Nanshan Ranch in Hunan has reached its best state in nearly a decade[N]. China Green Times, 2022-11-02(2) 李佑江. 湖南南山牧场草相呈现近10年最好景况[N]. 中国绿色时报, 2022-11-02(2) WU S P. Relationship between biodiversity and ecosystem versatility of different degraded grasslands in Nanshan, Hunan Province[D]. Changsha:Central South University of Forestry & Technology, 2025:9-54 吴沈萍. 湖南南山不同退化草地生物多样性与生态系统多功能性的关系[D]. 长沙:中南林业科技大学, 2025:9-54 General Administration of Quality Supervision, Inspection and Quarantine of the People’s Republic of China. GB 19377-2003 Classification Indicators for Degradation, Desertification, and Salinization of Natural Grasslands: GB 19377-2003 [S]. Beijing: China Standards Press, 2004: 1-5 国家质量监督检验检疫总局. GB 19377-2003天然草地退化、沙化、盐渍化的分级指标:GB 19377—2003[S]. 北京:中国标准出版社, 2004: 1-5 HAO A H, XUE X, PENG F, et al. Different vegetation and soil degradation characteristics of a typical grassland in the Qinghai-Tibetan Plateau[J]. Acta Ecologica Sinica, 2020, 40(3): 964-975 郝爱华, 薛娴, 彭飞, 等. 青藏高原典型草地植被退化与土壤退化研究[J]. 生态学报, 2020, 40(3): 964-975 BAO S D. Soil agro-chemistry analysis[M]. 3rd. Beijing: China Agriculture Press, 2000:30-48 鲍士旦. 土壤农化分析[M]. 3版. 北京:中国农业出版社, 2000:30-48 INSTITUTE OF SOIL SCIENCE, CHINESE ACADEMY OF SCIENCES. Soil physical and chemical analysis [M]. Shanghai: Shanghai Scientific & Technical Publishers, 1978:514-518 中国科学院南京土壤研究所. 土壤理化分析[M]. 上海:上海科学技术出版社, 1978:514-518 于文竹, 魏霞, 赵恒策, 等. 青藏高原高寒草原草甸土壤团聚 体及养分因子变化特征[J]. 水土保持学报, 2020, 34(6): 301-308, 317 YU W Z, WEI X, ZHAO H C, et al. Changes of Soil Aggregates and Nutrient Factors in Alpine Grassland and Alpine meadow of Qinghai Tibet Plateau[J]. Journal of Soil and Water Conservation, 2020, 34(6):301-308, 317 YANG P L, LUO Y P, SHI Y C. Fractal characteristics of soil characterized by weight distribution of particle size[J]. Cinese Science Bulletin, 1993, 38(20): 1896-1899 杨培岭, 罗远培, 石元春. 用粒径的重量分布表征的土壤分形特征[J]. 科学通报, 1993, 38(20): 1896-1899 HAI L, YAO T, ZHANG W M, et al. Distribution characteristics of soil aggregates and its organic carbon of alfalfa grassland with different growth years in Loess Hilly-gully region[J]. Grassland and Turf, 2020, 40(4): 16-21 海龙, 姚拓, 张文明, 等. 黄土丘陵沟壑区不同种植年限紫花苜蓿草地土壤团聚体及其有机碳分布特征[J]. 草原与草坪, 2020, 40 (4): 16-21 HAMZA A, KARČAUSKIENĖ D, MOCKEVIČIENĖ I, et al. Soil Aggregate Dynamics and Stability: Natural and Anthropogenic Drivers[J]. Agriculture, 2025, 15(23): 2500 TOU C Y, LIU X, GAN J J, et al. Plant residue quality regulates its incorporation into soil aggregates and organic carbon stabilisation[J]. Geoderma, 2025, 460: 117407 LU Z J, WANG H S, LIU J Z, et al. From waste to soil: The pivotal role of organic matter in soil aggregate formation under dry and wet conditions[J]. Plant and Soil, 2025, 514(1): 1543-1559 VINDUŠKOVÁ O, DECKMYN G, REYNAERT S, et al. More persistent precipitation regimes induce soil degradation[J]. Geoderma, 2025, 455: 117230 LI H R, LIU G, LIU Y, et al. Variability of soil aggregate disintegration during splash erosion under different circumstances[J]. Land Degradation & Development, 2025, 36(2): 454-465 AL-JEZNAWI D, AL-JANABI M A Q, ALMEIDA BERNARDO L F, et al. Modeling and mechanisms of desiccation-induced cracking and curling in clay soils: a state-of-the-art review[J]. Geotechnical and Geological Engineering, 2025, 43(7): 342 TONG K W, PEREIRA J M, HUANG K, et al. Experimental study on the microscopic swelling-shrinking mechanisms of clay-sand mixtures from wetting to drying[J]. Acta Geotechnica, 2025, 20(7): 3477-3500 CHEN Y, LV X F, CAO L T, et al. Rainfall-and leakage-induced soil erosion and debonding-migration effects[J]. Journal of Mountain Science, 2025, 22(9): 3200-3213 PAN Z D, CAI X M, BO Y M, et al. Response of soil organic carbon and soil aggregate stability to changes in land use patterns on the Loess Plateau[J]. Scientific Reports, 2024, 14(1): 31775 LIU X, MA Z W, QIN W P, et al. Enhanced unprotected organic carbon in macroaggregates facilitates soil organic carbon accumulation following grazing removal in Tibetan Alpine meadows[J]. Plant and Soil, 2025: 1-19 CHEN Y L, KUZYAKOV Y, MA Q W, et al. Aggregate size mediates the stability and temperature sensitivity of soil organic carbon in response to decadal biochar and straw amendments[J]. Soil Biology and Biochemistry, 2025, 211: 109969 CHEN W J, CHEN K L, WU F Z. Macroaggregate disintegration mediates forest gap impacts on MAOC decrease in karst ecosystems[J]. Geoderma, 2026, 465: 117661 SUN H, KUMAR A, VAN ZWIETEN L, et al. Soil aggregates mediate the thermal response of soil carbon mineralization and microbial enzyme activity[J]. International Journal of Environmental Research, 2025, 19(4): 106 BAI L L, DUAN J X, SHI P, et al. Erosion-deposition processes drive soil organic carbon mineralization through aggregate breakdown and buildup[J]. Catena, 2025, 260: 109432 ABDELRHMAN A ALI, SAYED Y A, FADL M E, et al. Impact of aggregate-associated carbon on soil mechanical properties: stability and compaction indices in pomegranate orchards of different ages[J]. Soil Systems, 2025, 9(1): 20 HAN Z X, WU X P, GAO H Z, et al. Long-term conservation tillage increases soil organic carbon stability by modulating microbial nutrient limitations and aggregate protection[J]. Agronomy, 2025, 15(7): 1571 WANG G Y, LI J Y, MAO J F, et al. Multiscale drivers and tipping points regulating particulate and mineral-associated organic carbon across Central Asian grasslands[J]. Geoderma, 2025, 464: 117610 HU M G, YAO Z Y, SHI L N, et al. Patchy degradation-induced changes in soil aggregates and organic carbon in an alpine swamp meadow on the Qinghai-Tibetan Plateau[J]. Land Degradation & Development, 2024, 35(16): 4713-4725)
Structural Regulation Mechanisms of Soil Aggregate Reorganization and Carbon Pool Responses to Grassland Degradation in Humid Regions
HUANG Hui-qun, LIU Xi-yuan, ZHU Xiao-hua, ZENG He-ping
Affiliations
1 Shaoyang Grassland Resources Protection Center, Shaoyang, Hunan Province 422000, China; 2 City college, Kunming University of Science and Technology, Kunming, Yunnan Province 650500, China
Grassland degradation is a major driver affecting soil structure stability and carbon cycling processes; however, its underlying pathways and regulatory mechanisms under humid climatic conditions remain insufficiently quantified. In this study, a typical montane meadow in Nanshan Pasture, Hunan Province, China, was selected to investigate the characteristics of soil aggregate structure and organic carbon distribution across a degradation gradient, including non-degraded (ND), lightly degraded (LD), moderately degraded (MD), and heavily degraded (HD) grasslands within the 0-30 cm soil layer. The results showed that, with increasing degradation intensity, the proportion of >5 mm macroaggregates decreased by approximately 47%, whereas the proportion of <0.25 mm microaggregates increased by approximately 168%. Meanwhile, the mean weight diameter (MWD) and geometric mean diameter (GMD) of both mechanically stable and water-stable aggregates declined by approximately 46% and 53%, respectively, accompanied by a significant increase in fractal dimension, indicating a transition of soil structure from a stable state to a fragmented state. The breakdown of macroaggregates released organic carbon, part of which was subsequently adsorbed and stabilized by microaggregates, resulting in a shift in carbon distribution from a “macroaggregate-dominated” pattern to a “microaggregate-dominated” pattern. Grassland degradation also led to a decline in total soil organic carbon and induced substantial structural reorganization of soil carbon pools, highlighting the pivotal role of soil aggregates in carbon protection and redistribution. These findings elucidate the structural mechanisms underlying carbon pool dynamics driven by grassland degradation in humid regions and provide a theoretical basis for ecological restoration and carbon sequestration management of degraded grasslands.
YAHDJIAN L, CAMPANA S, TOGNETTI P M, et al. Insights on global rangeland ecosystem services shaped by grazing and fertilization[J]. Frontiers in Ecology and the Environment, 2026, 24(3): e70022 WAHEED A, QIAO X, MUHAMMAD M, et al. Plant root-mediated carbon sequestration and nutrient cycling in grassland ecosystems under land use and climate change[J]. Agriculture, Ecosystems & Environment, 2025, 393: 109865 GUO H N, XIN X P, CHEN J Q, et al. Divergent responses of particulate and mineral-associated organic carbon stock to grazing in a Eurasian temperate meadow steppe[J]. Geoderma, 2025, 462: 117516 LI H N, MENG R B, MENG Z J, et al. Influence of grazing patterns on the stability of soil aggregates in semi-arid grasslands[J]. Journal of Arid Land, 2026, 18(2): 322-338 VAN DAEL T, VERWERFT Q, SMOLDERS E. The soil structural stability determined by the QuantiSlakeTest: Linkage with soil porosity, water-stable aggregate fractions and soil chemical properties[J]. Geoderma, 2025, 457: 117281 HU P L, ZHANG W, NOTTINGHAM A T, et al. Lithological controls on soil aggregates and minerals regulate microbial carbon use efficiency and necromass stability[J]. Environmental Science & Technology, 2024, 58(48): 21186-21199 QIN W P, YUAN H, SUN M Y, et al. Effects of Grazing Exclusion on Soil Aggregate Composition and Stability in Degraded Alpine Meadows[J/OL]. https://doi.org/10.19741/j.issn.1673-4831.2025.0554, 2025-10-29/2026-04-09 秦文萍, 袁晖, 孙铭阳, 等. 围栏封育对退化高寒草甸土壤团聚体组成及稳定性的影响[J/OL]. https://doi.org/10.19741/j.issn.1673-4831.2025.0554, 2025-10-29/2026-04-09 DU Z Y, CONG N, ZHAO G, et al. Divergent responses of vegetation and soil characteristics to grassland degradation in the Qinghai-Tibet and Inner Mongolia plateaus[J]. Catena, 2025, 257: 109146 LIN X, ZHANG S W, HUETE A, et al. Global-to-regional variations in the effects of grassland management on soil carbon[J]. Agriculture, Ecosystems & Environment, 2026, 400: 110251 ZHOU L, SU Z A, YANG J, et al. Effects of soil erosion on aggregate -associated organic carbon in typical grassland and terraced fields of the agro -pastoral ecotone of Qilian Mountains[J]. Journal of Soil and Water Conservation, 2025, 39(4): 198-207 周铃, 苏正安, 杨静, 等. 祁连山农牧交错带典型草地和梯田土壤侵蚀对团聚体有机碳的影响[J]. 水土保持学报, 2025, 39(4): 198-207 MA Q, BAI W, MOU G X, et al. Effects of warming on soil Aggregate stability and contents of carbon and nitrogen in Alpine swamp meadow [J]. Acta Agrestia Sinica, 2025, 33(2): 547-555 马群, 白炜, 牟国旭, 等. 增温对高寒沼泽草甸土壤团聚体稳定性及碳氮含量的影响[J]. 草地学报, 2025, 33(2): 547-555 MI J, WANG F, SHI J, et al. Contrasting trends in plant diversity and soil carbon mineralization under precipitation-driven vegetation and soil carbon dynamics in the Mongolian Plateau[J]. Ecology and Evolution, 2025, 15(7): e71806 LUO J Y. Driving mechanism of biochar combined enclosure on multifunctional restoration of degraded grassland ecosystem in Nanshan pasture[D]. Changsha:Central South University of Forestry & Technology, 2025:11 罗俊毅. 生物炭联合围封对南山牧场退化草地生态系统多功能恢复的驱动机制[D]. 长沙:中南林业科技大学, 2025:11 ZHANG W. Nanshan Grassland, where Laoshanjie is located[N]. China Cultural Relics News, 2023-02-21(1) 张伟. 老山界所在地——南山草原[N]. 中国文物报, 2023-02-21(1) YOU L S, LIU S M, XIAO T G, et al. Investigation on plant and insect species in Nanshan pasture farm, Hunan Province and insect fauna analysis[J].Journal of Hunan Agricultural University (Natural Sciences), 1999, 25(1): 47-51 游兰韶, 刘寿明, 肖铁光, 等. 湖南省南山牧场植物和昆虫种类调查及区系分析[J]. 湖南农业大学学报, 1999, 25(1): 47-51 ZHANG Y. Study on the changes and influencing factors of soil C∶N∶P in different grassland types in Nanshan, Hunan Province[D]. Changsha:Central South University of Forestry & Technology, 2023:9 张宇. 湖南南山不同草地类型土壤C∶N∶P变化及其影响因素研究[D]. 长沙:中南林业科技大学, 2023:9 LI Y J. The grassland condition of Nanshan Ranch in Hunan has reached its best state in nearly a decade[N]. China Green Times, 2022-11-02(2) 李佑江. 湖南南山牧场草相呈现近10年最好景况[N]. 中国绿色时报, 2022-11-02(2) WU S P. Relationship between biodiversity and ecosystem versatility of different degraded grasslands in Nanshan, Hunan Province[D]. Changsha:Central South University of Forestry & Technology, 2025:9-54 吴沈萍. 湖南南山不同退化草地生物多样性与生态系统多功能性的关系[D]. 长沙:中南林业科技大学, 2025:9-54 General Administration of Quality Supervision, Inspection and Quarantine of the People’s Republic of China. GB 19377-2003 Classification Indicators for Degradation, Desertification, and Salinization of Natural Grasslands: GB 19377-2003 [S]. Beijing: China Standards Press, 2004: 1-5 国家质量监督检验检疫总局. GB 19377-2003天然草地退化、沙化、盐渍化的分级指标:GB 19377—2003[S]. 北京:中国标准出版社, 2004: 1-5 HAO A H, XUE X, PENG F, et al. Different vegetation and soil degradation characteristics of a typical grassland in the Qinghai-Tibetan Plateau[J]. Acta Ecologica Sinica, 2020, 40(3): 964-975 郝爱华, 薛娴, 彭飞, 等. 青藏高原典型草地植被退化与土壤退化研究[J]. 生态学报, 2020, 40(3): 964-975 BAO S D. Soil agro-chemistry analysis[M]. 3rd. Beijing: China Agriculture Press, 2000:30-48 鲍士旦. 土壤农化分析[M]. 3版. 北京:中国农业出版社, 2000:30-48 INSTITUTE OF SOIL SCIENCE, CHINESE ACADEMY OF SCIENCES. Soil physical and chemical analysis [M]. Shanghai: Shanghai Scientific & Technical Publishers, 1978:514-518 中国科学院南京土壤研究所. 土壤理化分析[M]. 上海:上海科学技术出版社, 1978:514-518 于文竹, 魏霞, 赵恒策, 等. 青藏高原高寒草原草甸土壤团聚 体及养分因子变化特征[J]. 水土保持学报, 2020, 34(6): 301-308, 317 YU W Z, WEI X, ZHAO H C, et al. Changes of Soil Aggregates and Nutrient Factors in Alpine Grassland and Alpine meadow of Qinghai Tibet Plateau[J]. Journal of Soil and Water Conservation, 2020, 34(6):301-308, 317 YANG P L, LUO Y P, SHI Y C. Fractal characteristics of soil characterized by weight distribution of particle size[J]. Cinese Science Bulletin, 1993, 38(20): 1896-1899 杨培岭, 罗远培, 石元春. 用粒径的重量分布表征的土壤分形特征[J]. 科学通报, 1993, 38(20): 1896-1899 HAI L, YAO T, ZHANG W M, et al. Distribution characteristics of soil aggregates and its organic carbon of alfalfa grassland with different growth years in Loess Hilly-gully region[J]. Grassland and Turf, 2020, 40(4): 16-21 海龙, 姚拓, 张文明, 等. 黄土丘陵沟壑区不同种植年限紫花苜蓿草地土壤团聚体及其有机碳分布特征[J]. 草原与草坪, 2020, 40 (4): 16-21 HAMZA A, KARČAUSKIENĖ D, MOCKEVIČIENĖ I, et al. Soil Aggregate Dynamics and Stability: Natural and Anthropogenic Drivers[J]. Agriculture, 2025, 15(23): 2500 TOU C Y, LIU X, GAN J J, et al. Plant residue quality regulates its incorporation into soil aggregates and organic carbon stabilisation[J]. Geoderma, 2025, 460: 117407 LU Z J, WANG H S, LIU J Z, et al. From waste to soil: The pivotal role of organic matter in soil aggregate formation under dry and wet conditions[J]. Plant and Soil, 2025, 514(1): 1543-1559 VINDUŠKOVÁ O, DECKMYN G, REYNAERT S, et al. More persistent precipitation regimes induce soil degradation[J]. Geoderma, 2025, 455: 117230 LI H R, LIU G, LIU Y, et al. Variability of soil aggregate disintegration during splash erosion under different circumstances[J]. Land Degradation & Development, 2025, 36(2): 454-465 AL-JEZNAWI D, AL-JANABI M A Q, ALMEIDA BERNARDO L F, et al. Modeling and mechanisms of desiccation-induced cracking and curling in clay soils: a state-of-the-art review[J]. Geotechnical and Geological Engineering, 2025, 43(7): 342 TONG K W, PEREIRA J M, HUANG K, et al. Experimental study on the microscopic swelling-shrinking mechanisms of clay-sand mixtures from wetting to drying[J]. Acta Geotechnica, 2025, 20(7): 3477-3500 CHEN Y, LV X F, CAO L T, et al. Rainfall-and leakage-induced soil erosion and debonding-migration effects[J]. Journal of Mountain Science, 2025, 22(9): 3200-3213 PAN Z D, CAI X M, BO Y M, et al. Response of soil organic carbon and soil aggregate stability to changes in land use patterns on the Loess Plateau[J]. Scientific Reports, 2024, 14(1): 31775 LIU X, MA Z W, QIN W P, et al. Enhanced unprotected organic carbon in macroaggregates facilitates soil organic carbon accumulation following grazing removal in Tibetan Alpine meadows[J]. Plant and Soil, 2025: 1-19 CHEN Y L, KUZYAKOV Y, MA Q W, et al. Aggregate size mediates the stability and temperature sensitivity of soil organic carbon in response to decadal biochar and straw amendments[J]. Soil Biology and Biochemistry, 2025, 211: 109969 CHEN W J, CHEN K L, WU F Z. Macroaggregate disintegration mediates forest gap impacts on MAOC decrease in karst ecosystems[J]. Geoderma, 2026, 465: 117661 SUN H, KUMAR A, VAN ZWIETEN L, et al. Soil aggregates mediate the thermal response of soil carbon mineralization and microbial enzyme activity[J]. International Journal of Environmental Research, 2025, 19(4): 106 BAI L L, DUAN J X, SHI P, et al. Erosion-deposition processes drive soil organic carbon mineralization through aggregate breakdown and buildup[J]. Catena, 2025, 260: 109432 ABDELRHMAN A ALI, SAYED Y A, FADL M E, et al. Impact of aggregate-associated carbon on soil mechanical properties: stability and compaction indices in pomegranate orchards of different ages[J]. Soil Systems, 2025, 9(1): 20 HAN Z X, WU X P, GAO H Z, et al. Long-term conservation tillage increases soil organic carbon stability by modulating microbial nutrient limitations and aggregate protection[J]. Agronomy, 2025, 15(7): 1571 WANG G Y, LI J Y, MAO J F, et al. Multiscale drivers and tipping points regulating particulate and mineral-associated organic carbon across Central Asian grasslands[J]. Geoderma, 2025, 464: 117610 HU M G, YAO Z Y, SHI L N, et al. Patchy degradation-induced changes in soil aggregates and organic carbon in an alpine swamp meadow on the Qinghai-Tibetan Plateau[J]. Land Degradation & Development, 2024, 35(16): 4713-4725