Article(id=1309573314708857271, tenantId=1146029695717560320, journalId=1308397196074385509, issueId=1309573278520406630, articleNumber=null, orderNo=null, doi=10.11733/j.issn.1007-0435.2026.05.024, pmid=null, cstr=null, oa=null, hot=0, price=null, onlineType=0, articleFormat=0, articleType=null, articleTypeStr=null, receivedDate=1756569600000, receivedDateStr=2025-08-31, revisedDate=1766505600000, revisedDateStr=2025-12-24, acceptedDate=null, acceptedDateStr=null, onlineDate=1790156208922, onlineDateStr=2026-09-23, pubDate=null, pubDateStr=null, doiRegisterDate=null, doiRegisterDateStr=null, onlineIssueDate=1790156208922, onlineIssueDateStr=2026-09-23, onlineJustAcceptDate=null, onlineJustAcceptDateStr=null, onlineFirstDate=null, onlineFirstDateStr=null, sourceXml=null, magXml=null, createTime=1790156208922, creator=13701087609, updateTime=1790156208922, updator=13701087609, issue=Issue{id=1309573278520406630, tenantId=1146029695717560320, journalId=1308397196074385509, year='2026', volume='34', issue='5', pageStart='1557', pageEnd='1962', issueExtLink='null', onlineDate='null', pubDate='1778774400000', pubDateStr='2026-05-15', beforeIssueId=null, nextIssueId=null, price=null, status=1, issueComplete=1, articleOrder=1, issueType=-1, specialIssue=null, createTime=1790156200293, creator='13701087609', updateTime=1790156821973, updator='13701087609', preIssue=null, nextIssue=null, articleTotal=null, ext={EN=IssueExt(id=1309575886110814336, tenantId=1146029695717560320, journalId=1308397196074385509, issueId=1309573278520406630, language=EN, specialIssueTitle=, coverIllustrator=null, specialIssueEditor=, specialIssueAbout=), CN=IssueExt(id=1309575886115008641, tenantId=1146029695717560320, journalId=1308397196074385509, issueId=1309573278520406630, language=CN, specialIssueTitle=, coverIllustrator=null, specialIssueEditor=, specialIssueAbout=)}, issueFiles=null, downloadFileDto=null}, startPage=1827, endPage=1840, ext={EN=ArticleExt(id=1309573315027624378, articleId=1309573314708857271, tenantId=1146029695717560320, journalId=1308397196074385509, language=EN, title=The Influence of Grazing and Topography on the Main Populations and Population-Community Relationship of Alpine Shrublands in the Qilian Mountains, columnId=null, journalTitle=Acta Agrestia Sinica, columnName=null, runingTitle=null, highlight=null, articleAbstract=Grazing and topography are key factors influencing the structure and function of alpine shrub ecosystems, but their effects on shrub populations and communities remain unclear. For this purpose, this study explored the effects of topography (three types of slope positions) and grazing (three distances from livestock enclosure) on the dominant species of alpine shrubland Caragana jubata and the companion species Thalictrum alpinum in the Qilian Mountain Grassland Agricultural Demonstration Area. The results showed that: 1) The distance from livestock enclosure explained the plant height and height-to-crown ratio of the Caragana jubata as 45.10% and 46.80% (P<0.01), and the plant height, height-to-crown ratio and density of the Thalictrum alpinum as 46.50%, 40.60%(P<0.01) and 55.30% (P<0.001). The topography only affects the population coverage of the Caragana jubata, which can explain that the coverage of the Caragana jubata is 22.30% (P<0.05). 2) The overlap degree of the ecological niches of the Caragana jubata and the Thalictrum alpinum is the lowest at a distance of 400 meters between the livestock enclosure and the highest at 1200 meters. 3) The plant height, crown breadth and density of the population of Thalictrum alpinum significantly affected the community coverage, with an interpretation rate of 35.3% (P<0.05). The community coverage significantly affected the population density of Thalictrum alpinum, with an interpretation rate of 25.1%(P<0.01). The plant height, density and importance value of the population of Caragana jubata significantly affected the community coverage, with an interpretation rate of 57.6%(P<0.01). This study reveals the role of grazing and topography in influencing the shrub-herbaceous competitive balance in alpine shrublands by regulating population growth and its population-community relationship, providing a scientific basis for grazing in complex topography., authors=PENG Bai-qing1, SANG Ya-zhuan1, NING Jiao1, HOU Fu-jiang1, authorsList=PENG Bai-qing, SANG Ya-zhuan, NING Jiao, HOU Fu-jiang, authorCompany=1 China-Kazakhstan Belt and Road Joint Laboratory on Grassland Ecological Restoration, Key Laboratory of Grassland Livestock Industry Innovation, Ministry of Agriculture and Rural Affairs, Engineering Technology Research Center for Ecological Restoration and Utilization of Degraded Grassland in Northwest China, National Forestry and Grassland Administration, College of Grassland Agricultural Science and Technology, Lanzhou University, Lanzhou, Gansu Province 730020, China, correspAuthors=null, authorNote=null, correspAuthorsNote=null, copyrightStatement=null, copyrightOwner=null, extLink=null, articleAbsUrl=null, sourceXml=null, magXml=null, pdfUrl=null, pdf=null, pdfFileSize=null, pdfExtLink=null, richHtmlUrl=null, mobilePdfUrl=null, reviewReport=null, pdfFirstPage=null, abstractGraph=null, abstractGraphContent=null, abstractVideo=null, citation=null, cebUrl=null, magXmlContent=null, mapNumber=null, fund=null), CN=ArticleExt(id=1309573314960515513, articleId=1309573314708857271, tenantId=1146029695717560320, journalId=1308397196074385509, language=CN, title=放牧和地形对祁连山高寒灌丛主要种群及种群-群落关系的影响, columnId=1309565024058835950, journalTitle=草地学报, columnName=研究论文, runingTitle=null, highlight=null, articleAbstract=放牧和地形是影响高寒灌丛生态系统结构与功能的关键因素,但二者对灌丛种群及群落的作用尚不清楚。为此,本研究在祁连山草地农业示范区探究地形(3种坡位)与放牧(3个畜圈距离)对高寒灌丛优势种鬼箭锦鸡儿Caragana jubata和伴生种高山唐松草Thalictrum alpinum的影响,结果表明:(1)畜圈距离解释鬼箭锦鸡儿株高和株高冠幅比45.10%和46.80%(P<0.01),解释高山唐松草株高、株高冠幅比和密度46.50%,40.60%(P<0.01)和55.30%(P<0.001);地形仅影响鬼箭锦鸡儿种群盖度,可解释鬼箭锦鸡儿盖度22.30%(P<0.05);(2)鬼箭锦鸡儿和高山唐松草生态位重叠度在畜圈距离400 m时最低,1200 m时最高;(3)高山唐松草的种群的株高、冠幅和密度显著影响群落盖度,解释率为35.3%(P<0.05);群落盖度显著影响高山唐松草种群密度,解释率为25.1%(P<0.01),鬼箭锦鸡儿种群的株高、密度和重要值显著影响群落盖度,解释率为57.6%(P<0.01)。本研究揭示了放牧和地形通过调控种群生长及其种群-群落关系,影响高寒灌丛中灌木-草本竞争平衡的作用,为复杂地形下的放牧提供科学依据。, authors=彭白青1, 桑亚转1, 宁娇1, 侯扶江1, authorsList=彭白青, 桑亚转, 宁娇, 侯扶江, authorCompany=1 中国-哈萨克斯坦草原生态修复"一带一路"联合实验室/农业农村部草牧业创新重点实验室/国家林业草原局西北退化草原生态修复与利用工程技术研究中心/兰州大学草地农业科技学院, 甘肃 兰州 730020, correspAuthors=null, authorNote=彭白青(2002-),女,汉族,甘肃永登人,硕士研究生,主要从事草地生态方向研究,E-mail:pengbq2024@lzu.edu.cn, correspAuthorsNote=null, copyrightStatement=null, copyrightOwner=null, extLink=null, articleAbsUrl=null, sourceXml=null, magXml=null, pdfUrl=null, pdf=quwlryqR5VAMlymqnB1jCw==, pdfFileSize=2126829, pdfExtLink=null, richHtmlUrl=null, mobilePdfUrl=null, reviewReport=null, pdfFirstPage=null, abstractGraph=null, abstractGraphContent=null, abstractVideo=null, citation=null, cebUrl=null, magXmlContent=null, mapNumber=null, fund=国家重点研发计划 (2025YFE0212000); 甘肃省拔尖领军人才 (GSBJLJ-2022-20); 甘肃省创新平台计划项目 (26JDWA001))}, authors=null, keywords=[Keyword(id=1309573315161842107, tenantId=1146029695717560320, journalId=1308397196074385509, articleId=1309573314708857271, language=CN, orderNo=1, keyword=种群), Keyword(id=1309573315233145276, tenantId=1146029695717560320, journalId=1308397196074385509, articleId=1309573314708857271, language=CN, orderNo=2, keyword=群落), Keyword(id=1309573315304448445, tenantId=1146029695717560320, journalId=1308397196074385509, articleId=1309573314708857271, language=CN, orderNo=3, keyword=株高), Keyword(id=1309573315392528830, tenantId=1146029695717560320, journalId=1308397196074385509, articleId=1309573314708857271, language=CN, orderNo=4, keyword=冠幅), Keyword(id=1309573315451249087, tenantId=1146029695717560320, journalId=1308397196074385509, articleId=1309573314708857271, language=CN, orderNo=5, keyword=伴生种), Keyword(id=1309573315539329472, tenantId=1146029695717560320, journalId=1308397196074385509, articleId=1309573314708857271, language=EN, orderNo=1, keyword=Population), Keyword(id=1309573315614826945, tenantId=1146029695717560320, journalId=1308397196074385509, articleId=1309573314708857271, language=EN, orderNo=2, keyword=Community), Keyword(id=1309573315686130114, tenantId=1146029695717560320, journalId=1308397196074385509, articleId=1309573314708857271, language=EN, orderNo=3, keyword=Height), Keyword(id=1309573315761627587, tenantId=1146029695717560320, journalId=1308397196074385509, articleId=1309573314708857271, language=EN, orderNo=4, keyword=Crown breadth), Keyword(id=1309573315832930756, tenantId=1146029695717560320, journalId=1308397196074385509, articleId=1309573314708857271, language=EN, orderNo=5, keyword=Companion species)], refs=null, funds=null, companyList=null, figs=null, attaches=null, journal=Journal(id=1308395841968504927, delFlag=0, nameCn=草地学报, nameEn=Acta Agrestia Sinica, nameHistory1=null, nameHistory2=null, issn=1007-0435, eissn=null, cn=11-3362/S, coden=null, periodic=0, language=CN, oaType=null, ccby=null, superviseOffice=null, ownerOffice=null, pubOffice=null, editorOffice=null, officeType=null, aims=null, clcCode=null, officeProv=null, officeCity=null, officeAddr=null, officeZip=null, officeEmail=null, officePhone=null, editDirector=null, officeDirector=null, officeDirectorPhone=null, officeStaffNum=null, officeEmpNum=null, coverPicUrl=gCdi9sXCX82eLR6S1d4VLg==, journalPrice=null, startedYear=null, abbrevIsoEn=Acta Agrestia Sinica, journalRemark=null, publicationField=null, createdTime=1789875477545, updatedTime=1789876241473, createdBy=18614031015, updatedBy=13701087609, firstLetterCn=C, firstLetterEn=C, subjectCode=Agriculture, subjectName=null, subjectCodeEn=Agriculture, subjectNameEn=null, picCn=gCdi9sXCX82eLR6S1d4VLg==, picEn=4dKqEDfsaGuyf+bpm/3gfw==, jcr=null, cjcr=null, exts=[JournalExt(id=1308399046192812446, language=CN, name=草地学报, nameHistory1=null, nameHistory2=null, managedBy=, sponsoredBy=, publishedBy=, editorOffice=, officeProv=null, officeCity=null, officeAddr=, officeZip=, editDirector=, officeDirector=null, officePhone=null, coverPicUrl=null, journalRemark=, submitArticleUrl=null, websiteUrl=, createdTime=1789876241491, updatedTime=1789876241491, createdBy=13701087609, updatedBy=13701087609, submissionGuidelinesUrl=, submissionAuthorUrl=https://manu40.magtech.com.cn/JournalX_cdxb/authorLogOn.action, submissionEditorUrl=https://manu40.magtech.com.cn/JournalX_cdxb/editorLogOn.action, submissionReviewUrl=https://manu40.magtech.com.cn/JournalX_cdxb/expertLogOn.action, submissionCeEditorUrl=, submissionAeEditorUrl=, option={"copyright":""}), JournalExt(id=1308399046268309919, language=EN, name=Acta Agrestia Sinica, nameHistory1=null, nameHistory2=null, managedBy=, sponsoredBy=, publishedBy=, editorOffice=, officeProv=null, officeCity=null, officeAddr=, officeZip=, editDirector=, officeDirector=null, officePhone=null, coverPicUrl=null, journalRemark=, submitArticleUrl=null, websiteUrl=, createdTime=1789876241509, updatedTime=1789876241509, createdBy=13701087609, updatedBy=13701087609, submissionGuidelinesUrl=, submissionAuthorUrl=https://manu40.magtech.com.cn/JournalX_cdxb/authorLogOn.action, submissionEditorUrl=https://manu40.magtech.com.cn/JournalX_cdxb/editorLogOn.action, submissionReviewUrl=https://manu40.magtech.com.cn/JournalX_cdxb/expertLogOn.action, submissionCeEditorUrl=, submissionAeEditorUrl=, option={"copyright":""})], databaseList=null, tenantJournalId=1308397196074385509, websiteList=[Website(id=1308429426430071695, webName=null, webTitle=null, webDomain=null, webCopyrigh=null, webIpcNo=null, seoTitle=null, seoKeywords=null, seoDescription=null, tenantJournalId=null, journalId=1308397196074385509, journalNameCn=null, journalNameEn=null, grayFlag=null, tenantId=1146029695717560320, platformId=null, journalGroupId=null, journalGroupNameCn=null, journalGroupNameEn=null, type=1, domain=https://castjournals.cast.org.cn/joweb/cdxb/CN, language=CN, createTime=1789883484704, createBy=18614031015, updateTime=1789883745058, updateBy=18614031015, name=草地学报-中文, tplId=1146099689490845704, title=草地学报, delFlag=0, indexPage=/home, props=[WebsiteProps(id=1308430669353340972, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1308429426430071695, code=articleTextType, value=kx, createTime=1789883781040, updateTime=1789883781040, creator=18614031015, updator=18614031015), WebsiteProps(id=1308430669328175145, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1308429426430071695, code=banner, value=null, createTime=1789883781034, updateTime=1789883781034, creator=18614031015, updator=18614031015), WebsiteProps(id=1308430669382701103, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1308429426430071695, code=grayFlag, value=0, createTime=1789883781047, updateTime=1789883781047, creator=18614031015, updator=18614031015), WebsiteProps(id=1308430669319786536, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1308429426430071695, code=logo, value=https://castjournals.cast.org.cn/joweb/cdxb/CN/file/pic?fileId=BZ28lIoKdB54MmA0z6K66g==, createTime=1789883781032, updateTime=1789883781032, creator=18614031015, updator=18614031015), WebsiteProps(id=1308430669395284017, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1308429426430071695, code=minRunFlag, value=0, createTime=1789883781050, updateTime=1789883781050, creator=18614031015, updator=18614031015), WebsiteProps(id=1308430669344952363, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1308429426430071695, code=picServerUrl, value=https://castjournals.cast.org.cn/joweb/cdxb/CN/file/pic, createTime=1789883781038, updateTime=1789883781038, creator=18614031015, updator=18614031015), WebsiteProps(id=1308430669391089712, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1308429426430071695, code=silenceFlag, value=0, createTime=1789883781049, updateTime=1789883781049, creator=18614031015, updator=18614031015), WebsiteProps(id=1308430669336563754, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1308429426430071695, code=staticResourcePath, value=https://castjournals.cast.org.cn/joweb/cast_kjdb_cn_619/, createTime=1789883781036, updateTime=1789883781036, creator=18614031015, updator=18614031015), WebsiteProps(id=1308430669361729581, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1308429426430071695, code=themeColor, value=null, createTime=1789883781042, updateTime=1789883781042, creator=18614031015, updator=18614031015), WebsiteProps(id=1308430669370118190, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1308429426430071695, code=themeStyle, value=null, createTime=1789883781044, updateTime=1789883781044, creator=18614031015, updator=18614031015)]), Website(id=1308429426501374870, webName=null, webTitle=null, webDomain=null, webCopyrigh=null, webIpcNo=null, seoTitle=null, seoKeywords=null, seoDescription=null, tenantJournalId=null, journalId=1308397196074385509, journalNameCn=null, journalNameEn=null, grayFlag=null, tenantId=1146029695717560320, platformId=null, journalGroupId=null, journalGroupNameCn=null, journalGroupNameEn=null, type=1, domain=https://castjournals.cast.org.cn/joweb/cdxb/EN, language=EN, createTime=1789883484721, createBy=18614031015, updateTime=1789883716139, updateBy=18614031015, name=草地学报-英文, tplId=1146101810881728533, title=Acta Agrestia Sinica, delFlag=0, indexPage=/home, props=[WebsiteProps(id=1308430635979264030, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1308429426501374870, code=articleTextType, value=kx, createTime=1789883773083, updateTime=1789883773083, creator=18614031015, updator=18614031015), WebsiteProps(id=1308430635954098203, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1308429426501374870, code=banner, value=null, createTime=1789883773077, updateTime=1789883773077, creator=18614031015, updator=18614031015), WebsiteProps(id=1308430636000235553, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1308429426501374870, code=grayFlag, value=0, createTime=1789883773088, updateTime=1789883773088, creator=18614031015, updator=18614031015), WebsiteProps(id=1308430635945709594, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1308429426501374870, code=logo, value=https://castjournals.cast.org.cn/joweb/cdxb/EN/file/pic?fileId=BZ28lIoKdB54MmA0z6K66g==, createTime=1789883773075, updateTime=1789883773075, creator=18614031015, updator=18614031015), WebsiteProps(id=1308430636012818467, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1308429426501374870, code=minRunFlag, value=0, createTime=1789883773091, updateTime=1789883773091, creator=18614031015, updator=18614031015), WebsiteProps(id=1308430635970875421, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1308429426501374870, code=picServerUrl, value=https://castjournals.cast.org.cn/joweb/cdxb/EN/file/pic, createTime=1789883773081, updateTime=1789883773081, creator=18614031015, updator=18614031015), WebsiteProps(id=1308430636004429858, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1308429426501374870, code=silenceFlag, value=0, createTime=1789883773089, updateTime=1789883773089, creator=18614031015, updator=18614031015), WebsiteProps(id=1308430635962486812, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1308429426501374870, code=staticResourcePath, value=https://castjournals.cast.org.cn/joweb/cast_kjdb_en_623/, createTime=1789883773079, updateTime=1789883773079, creator=18614031015, updator=18614031015), WebsiteProps(id=1308430635987652639, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1308429426501374870, code=themeColor, value=null, createTime=1789883773085, updateTime=1789883773085, creator=18614031015, updator=18614031015), WebsiteProps(id=1308430635991846944, tenantId=1146029695717560320, journalId=null, journalGroupId=null, siteId=1308429426501374870, code=themeStyle, value=null, createTime=1789883773086, updateTime=1789883773086, creator=18614031015, updator=18614031015)])], journalTitle=草地学报, weixinUrl=null, journalUrl=https://manu40.magtech.com.cn/Jweb_cdxb, iacademicId=null, status=1, seqNo=null, journalTitleEn=Acta Agrestia Sinica, journalPhotoCn=gCdi9sXCX82eLR6S1d4VLg==, journalPhotoEn=4dKqEDfsaGuyf+bpm/3gfw==, journalFirstLetter=C, journalRecommend=null, journalNew=null, journalCollection=null, jcrJf=null, cjcrJf=null, jcrJfStr=null, cjcrJfStr=null, submissionFirstDecision=null, sciSubjectClassification=null, casSubjectClassification=null, citeScore=null, totalCitationFrequency=null, icpCode=null, psCode=null, advertisingLicenseCode=null, copyrightInformation=null, country=null, option=, provinceCode=null, provinceName=null, collectFlag=false, interPubPlatform=, interPubPlatformUrl=null), detailUrlCn=https://castjournals.cast.org.cn/joweb/cdxb/CN/10.11733/j.issn.1007-0435.2026.05.024, detailUrlEn=https://castjournals.cast.org.cn/joweb/cdxb/EN/10.11733/j.issn.1007-0435.2026.05.024, pdfUrlCn=https://castjournals.cast.org.cn/joweb/cdxb/CN/PDF/10.11733/j.issn.1007-0435.2026.05.024, pdfUrlEn=https://castjournals.cast.org.cn/joweb/cdxb/EN/PDF/10.11733/j.issn.1007-0435.2026.05.024, aliStartDate=null, aliEndDate=null, collectionFlag=false, citedCount=null, citedUrl=null, previewStatus=0, delFlag=0, hasFullText=0, orderTime=1778774400000, fullTextJson=null, articleText=null, reference=HOU F J, YANG Z Y. Effects of grazing of livestock on grassland[J]. Acta Ecologica Sinica, 2006, 26(1):244-264 侯扶江, 杨中艺. 放牧对草地的作用[J]. 生态学报, 2006, 26(1):244-264 SONG W J, SU J S, ZHANG M D, et al. Plant compensatory growth and optimal grazing intensity of grasslands in northern China: A meta-analysis of grazing experiments[J]. Chinese Science Bulletin, 2023, 68(11):1330-1342 宋伟江, 苏纪帅, 张梦迪, 等. 中国北方草地植物补偿性生长与合理放牧强度:基于放牧实验的整合分析[J]. 科学通报, 2023, 68(11):1330-1342 JAMES M B, JOE F, MARK J S, et al.A plant trait analysis of responses to grazing in a long‐term experiment[J].Journal of Applied ecology, 2001, 38(2):253-267 FEI X, SUO C X, XIANG S, et al. Responses of plant community structure and functional group characteristics to long-term seasonal grazing in an alpine meadow on eastern Qinghai-Tibet Plateau[J]. Acta Agrestia Sinica, 2022, 30(8):1954-1963 费璇, 锁才序, 向双, 等. 青藏东缘高寒草甸植物群落结构及功能群特征对长期季节放牧的响应[J]. 草地学报, 2022, 30(8):1954-1963 DUAN H C, QI Y, KANG W P, et al. Seasonal variation of vegetation and its spatiotemporal response to climatic factors in the Qilian Mountains, China[J]. Sustainability, 2022, 14(9):4926 JING Y Y, XU C L, YU X J. The effect of grazing yak’s activity range on vegetation community structure and soil physicochemical characteristics in cold season pasture of alpine meadow[J]. Grassland and Turf, 2024, 44(4):54-63 景媛媛, 徐长林, 鱼小军. 牦牛放牧活动范围对高寒草甸冷季牧场植被群落结构和土壤理化性质的影响[J]. 草原与草坪, 2024, 44(4):54-63 BIAN J X, NIU X X, CHEN J, et al. Nest circle distance affects plant community structure and spatial distribution of soil nutrients in alpine meadows[J]. Pratacultural Science, 2024, 41(9):2003-2011 边金霞, 牛小霞, 陈娟, 等. 窝圈距离影响高寒草甸植物群落结构和土壤养分空间分布[J]. 草业科学, 2024, 41(9):2003-2011 SUN H, ZHAO Y G, LIU G L, et al. Effects of grazing intensity on the grassland vegetation community characteristics in the hilly Loess Plateau Region[J]. Acta Agrestia Sinica, 2022, 30(4):810-817 孙会, 赵允格, 刘广亮, 等. 放牧强度对黄土丘陵区草地植物群落特征的影响[J]. 草地学报, 2022, 30(4):810-817 WANG R H, PENG Q, ZHANG W D, et al. Ecohydrological service characteristics of Qilian Mountain ecosystem in the next 30 years based on scenario simulation[J]. Sustainability, 2022, 14(3):1819 ZHAO L P, BAI X, TAN S T, et al. Effect of different enclosure durations on aboveground vegetation in typical steppe on Loess Plateau[J]. Pratacultural Science, 2018, 35(1):27-35 赵凌平, 白欣, 谭世图, 等. 不同年限封育对黄土高原典型草原地上植被的影响[J]. 草业科学, 2018, 35(1):27-35 WANG L, XU D M, ZHANG J J. Effects of enclosure on composition of plant community and species diversity of desert steppe[J]. Pratacultural Science, 2012, 29(10):1512-1516 王蕾, 许冬梅, 张晶晶. 封育对荒漠草原植物群落组成和物种多样性的影响[J]. 草业科学, 2012, 29(10):1512-1516 PAN X X, LIU B R, NIU S F, et al. Effects of enclosure years on community structure and species diversity of desert grassland[J]. Guizhou Agricultural Sciences, 2018(7):95-99 潘笑笑, 刘秉儒, 牛宋芳, 等.围封年限对荒漠草原群落结构及物种多样性的影响[J].贵州农业科学, 2018, 46(7):95-99 XU H W, WANG X K, QU Q, et al. Variations and factors characterizing ecological niches of species in a stable grassland plant community[J]. Ecological Indicators, 2021, 128:107846 ZHANG F , BENNETT J A , ZHANG B , et al.Cessation of grazing stabilizes productivity through effects on species asynchrony and stability of shrub/semi-shrub plants in arid grasslands[J].Agriculture, Ecosystems & Environment, 2023(348):108411 WANG W , KANG Y , LIU M , et al.Shrub encroachment in alpine meadows facilitates soil carbon and nitrogen storage by redistributing soil water[J].Catena, 2025(256):109061 WANG L, MOU P, WANG T M. Impacts of topography on spatial variability of temperate understory biomass in closed forest: a case study of Northeast China Tiger and Leopard National Park[J]. Guihaia, 2024, 44(8):1512-1523 王乐, 牟溥, 王天明. 地形对温带密林下灌草层地上生物量分异的影响——以东北虎豹国家公园为例[J]. 广西植物, 2024, 44(8):1512-1523 LÜ Z L, LIU B, CHANG F, et al. Relationship between plant functional diversity and ecosystem multifunctionality in Bayanbulak alpine meadow along an altitude gradient[J]. Chinese Journal of Plant Ecology, 2023, 47(6):822-832 吕自立, 刘彬, 常凤, 等. 巴音布鲁克高寒草甸植物功能多样性与生态系统多功能性关系沿海拔梯度的变化[J]. 植物生态学报, 2023, 47(6):822-832 LI Q, RONG L, WANG M. Effects of topography on diversity and distribution pattern of plant species in karst mountains area[J]. Bulletin of Soil and Water Conservation, 2019, 39(6):27-34 李芹, 容丽, 王敏. 地形对喀斯特山地植物物种多样性及分布格局的影响[J]. 水土保持通报, 2019, 39(6):27-34 MA Y R, GUAN Q Y, SUN Y F, et al. Three-dimensional dynamic characteristics of vegetation and its response to climatic factors in the Qilian Mountains[J]. Catena, 2022, 208:105694 FENG X X, CHEN Q. Drivers grass-shrub competition in grassland ecosystems[J]. Chinese Journal of Grassland, 2024, 46(8):137-148 冯星星, 陈清. 草原生态系统影响草本-灌木竞争的主要因素[J]. 中国草地学报, 2024, 46(8):137-148 ZHANG P J, HUANG J H, MU L, et al. Influence of nitrogen and water addition on the primary productivity of Stipa breviflora in a desert steppe under different grazing intensities[J]. Acta Ecologica Sinica, 2022, 42(13):5458-5470 张璞进, 黄建辉, 木兰, 等. 氮水添加对放牧背景下荒漠草原生产力的影响[J]. 生态学报, 2022, 42(13):5458-5470 WANG J F, ZHANG L H, ZHAO R F, et al. Responses of plant growth of different life-forms to precipitation changes in desert steppe[J]. Chinese Journal of Applied Ecology, 2020, 31(3):778-786 王军锋, 张丽华, 赵锐锋, 等. 荒漠草原区不同生活型植物生长对降水变化的响应[J]. 应用生态学报, 2020, 31(3):778-786 GAO L X, LAN T Y, ZHAO Z X, et al. Spatial turnover of shrubland communities and underlying factors in northern mid-subtropical China[J]. Chinese Journal of Plant Ecology, 2022, 46(11):1411-1421 高璐鑫, 兰天元, 赵志霞, 等. 中国中亚热带北部灌丛群落植物空间周转及其驱动因素[J]. 植物生态学报, 2022, 46(11):1411-1421 WANG B B, ZHOU S Q, JING Y D, et al. Temporal-spatial variations of vegetation phenology and the influence of topography on phenology in Shanxi Province[J]. Chinese Journal of Ecology, 2021, 40(6):1839-1848 王贝贝, 周淑琴, 荆耀栋, 等. 山西省植被物候时空变化以及地形对物候的影响[J]. 生态学杂志, 2021, 40(6):1839-1848 XIAO X, GE R, HOU F J. Effects of grazing and topography on α diversity and physical properties of the soil of alpine meadow communities in the Qinghai-Tibet Plateau[J]. Pratacultural Science, 2019, 36(12):3041-3051 肖翔, 格日才旦, 侯扶江. 青藏高原放牧和地形对高寒草甸群落α多样性和土壤物理性质的影响[J]. 草业科学, 2019, 36(12):3041-3051 LI Y, ZHAO X R, LI G T, et al. Effect of grazing on soil microbial biomass phosphorus and phosphatase activities on different topographic unit of a typical steppe[J]. Acta Ecologica Sinica, 2022, 42(10):4137-4149 李营, 赵小蓉, 李贵桐, 等. 内蒙古典型草原不同地形单元放牧对土壤微生物量磷及磷酸酶活性的影响[J]. 生态学报, 2022, 42(10):4137-4149 HOWISON R A, OLFF H, VAN DE KOPPEL J, et al. Biotically driven vegetation mosaics in grazing ecosystems: the battle between bioturbation and biocompaction[J]. Ecological Monographs, 2017, 87(3):363-378 HOU S J, WANG Y X. Summer grazing of red deer effects on plant diversity of alpine grassland on Qilian Mountain[J]. Acta Prataculturae Sinica, 2020, 29(10):206-210 侯帅君, 王迎新. 马鹿夏季放牧对祁连山草原群落多样性的影响[J]. 草业学报, 2020, 29(10):206-210 REN J Z. Grassland Agroecology[M]. Beijing: China Agriculture Press, 1995: 51-84 任继周. 草地农业生态学[M]. 北京:中国农业出版社, 1995:51-84 LIU J X, GONG Y B, ZUO Q, et al.A review of mountain grazing-paths and vegetationspatial heterogeneity [J]. Acta Prataculturae Sinica, 2012, 21(1):254-261 刘金鑫, 宫渊波, 左琴, 等. 山地牧道与植被空间异质性研究评述[J]. 草业学报, 2012, 21(1):254-261 LU W D, ZHU H B. SPSS Statistical Analysis[M]. 5th ed. Beijing: Publishing House of Electronics Industry, 2015:286-301 卢纹岱, 朱红兵. SPSS统计分析[M]. 5版. 北京:电子工业出版社, 2015:286-301 SHIPLEY B. The AIC model selection method applied to path analytic models compared using a d-separation test[J]. Ecology, 2013, 94(3):560-564 ZHANG R, YU F Y, ZHOU R H, et al. Effects of slope position and aspect on structure and species diversity of shrub community in the Jiajin Mountains, Sichuan Province, China[J]. Chinese Journal of Applied Ecology, 2020, 31(8):2507-2514 张荣, 余飞燕, 周润惠, 等. 坡向和坡位对四川夹金山灌丛群落结构与物种多样性特征的影响[J]. 应用生态学报, 2020, 31(8):2507-2514 WANG Z H, LYU S J, WANG Z W, et al. Correlation analysis between the important value of dominant population and species diversity after spring rest grazing in desert steppe[J]. Chinese Journal of Grassland, 2025, 47(1):45-53 王梓晗, 吕世杰, 王忠武, 等. 荒漠草原春季休牧后优势种群重要值与物种多样性的关联性分析[J]. 中国草地学报, 2025, 47(1):45-53 QIN H, LI M G, GAO J, et al. Composition and diversity of soil bacterial communities in shrub at different altitudes in luya mountain[J]. Ecology and Environmental Sciences, 2023, 32(3):459-468 秦浩, 李蒙爱, 高劲, 等. 芦芽山不同海拔灌丛土壤细菌群落组成和多样性研究[J]. 生态环境学报, 2023, 32(3):459-468 FAN Y , LI X Y , HUANG H , et al.Does phenology play a role in the feedbacks underlying shrub encroachment?[J].Science of The Total Environment, 2018, 657(20):1064-1073 YANG W , QU G , KELLY A R , et al.Positive effects of leguminous shrub encroachment on multiple ecosystem functions of alpine meadows and steppes greatly depended on increasing soil nutrient[J].Catena, 2024(236):107745 LI Y F, CHEN L, LI X B, et al. Structural characteristics and ecological effects of a shrub-fertile island in a desert steppe[J]. Pratacultural Science, 2018, 35(10):2327-2335 李月飞, 陈林, 李学斌, 等. 荒漠草原灌丛沃岛的结构特征和生态效应[J]. 草业科学, 2018, 35(10):2327-2335 LIU Y Y, DING J Y. Research progress on shrub encroachment in China[J]. Chinese Journal of Grassland, 2025, 47(4):127-141 刘奕吟, 丁婧祎. 中国草地灌丛化研究进展[J]. 中国草地学报, 2025, 47(4):127-141 ZHANG X, YE M, CHEN W L, et al. Influence of topography on species diversity of grassland plant communities in the Altai Mountains[J]. Chinese Journal of Grassland, 2024, 46(7):57-71 张西, 叶茂, 陈维龙, 等. 地形对阿尔泰山草地植物群落物种多样性的影响[J]. 中国草地学报, 2024, 46(7):57-71 AKASH TARIQ, GAOY J, ZENG F J, et al. Guardians of arid lands: deep-rooted defense against desertification and climate change[J].Trends in Plant Science, 2026, 31(3):295-312 QIAO O M, CHEN Z. Plant diversity on different types of slope ecological engineering and its responses to environmental factors in mining areas[J]. Chinese Journal of Applied Ecology, 2022, 33(3):742-748 乔欧盟, 陈璋. 矿区不同类型生态护坡工程植物多样性对环境因子的响应[J]. 应用生态学报, 2022, 33(3):742-748 WANG W H, WANG S L, LIANG G L, et al. Effects of slope categories of differing aspect and position on plant community diversity in alpine shrubland in the Qilian Mountains[J]. Acta Prataculturae Sinica, 2025, 34(1):17-28 王文虎, 王世林, 梁国玲, 等. 坡向和坡位对祁连山高寒灌丛植物群落多样性的影响[J]. 草业学报, 2025, 34(1):17-28 SUN Y, HE M Z, WANG L. Effects of precipitation control on plant diversity and biomass in a desert region[J]. Acta Ecologica Sinica, 2018, 38(7):2425-2433 孙岩, 何明珠, 王立. 降水控制对荒漠植物群落物种多样性和生物量的影响[J]. 生态学报, 2018, 38(7):2425-2433 XIAO Y M, YANG L C, NIE X Q, et al. Effects of slope position on phylogenetic diversity and structure of alpine shrub community[J]. Chinese Journal of Ecology, 2019, 38(6):1611-1619 肖元明, 杨路存, 聂秀青, 等. 坡位对高寒灌丛群落谱系多样性和谱系结构的影响[J]. 生态学杂志, 2019, 38(6):1611-1619 CAO J Y, LIU J F, YUAN Q, et al. Traits of shrubs in forests and bushes reveal different life strategies[J]. Chinese Journal of Plant Ecology, 2020, 44(7):715-729 曹嘉瑜, 刘建峰, 袁泉, 等. 森林与灌丛的灌木性状揭示不同的生活策略[J]. 植物生态学报, 2020, 44(7):715-729 LIU Y Z, SUN C C, LIU W T, et al. Response of keystone species changes in alpine grassland plant communities to different herbivore assemblage grazing[J]. Acta Ecologica Sinica, 2022, 42(18):7529-7540 刘玉祯, 孙彩彩, 刘文亭, 等. 高寒草地植物群落关键种对不同放牧家畜组合放牧的响应[J]. 生态学报, 2022, 42(18):7529-7540 JIANG K W, ZHANG Q Q, WANG Y F, et al. Characteristics of plant functional groups and the relationships with soil environmental factors in middle part of northern slope of Tianshan Mountains under different grazing intensities[J]. Chinese Journal of Plant Ecology, 2024, 48(6):701-718 江康威, 张青青, 王亚菲, 等. 放牧干扰下天山北坡中段植物功能群特征及其与土壤环境因子的关系[J]. 植物生态学报, 2024, 48(6):701-718 HAN D Y, YANG Y F, ZHANG W, et al. Impact of cutting time and intensity simulating large herbivore grazing on compensatory growth of Elymus dahuricus[J]. Acta Ecologica Sinica, 2026, 46(1):582-592 韩大勇, 杨允菲, 张维, 等. 刈割强度与频率模拟大型草食动物采食对披碱草补偿生长的影响[J]. 生态学报, 2026, 46(1):582-592 ZHANG Y F, YANG X X, DONG Q M, et al. Effects of Mixed Grazing of Yak and Tibetan Sheep on Feed Intake of Grazing Livestock and Plant Compensation Growth[J]. Acta Agrestia Sinica, 2019, 27(6):1607-1614 张艳芬, 杨晓霞, 董全民, 等.牦牛和藏羊混合放牧对放牧家畜采食量和植物补偿性生长的影响[J].草地学报, 2019, 27(6):1607-1614 YANG Y, LIU A J, LI L H, et al. Effects of different disturbance types on plant species composition and functional group characteristics of typical steppe in Inner Mongolia, China[J]. Chinese Journal of Applied Ecology, 2016, 27(3):794-802 杨勇, 刘爱军, 李兰花, 等. 不同干扰方式对内蒙古典型草原植物种组成和功能群特征的影响[J]. 应用生态学报, 2016, 27(3):794-802 CANALS R M, MÚGICA L, DURÁN M, et al. Restorative pyric herbivory practices in shrub-encroached grasslands enhance nutrient resource availability and spatial heterogeneity[J]. Agriculture, Ecosystems & Environment, 2024(372):109072 MI Y Q, XU H B, ZHANG L, et al. The impact of enclosure on vegetation ecological niche and interspecific linkage in Inner Mongolian desert steppe[J]. Acta Agrestia Sinica2026, 34(3):1007-1016 米雨晴, 许宏斌, 张雷, 等. 围封对内蒙古荒漠草原植被生态位及种间联结性的影响[J]. 草地学报, 2026, 34(3):1007-1016 LI L, LI D, MHKNOPS J, et al. Assessing grazing intensity at pasture and quadrat scales for sustainable pastoralism on the Qinghai-Tibetan Plateau[J]. Acta Ecologica Sinica, 2025, 45(10):4952-4962 李黎, 李丹, MHKNOPS Johannes, 等. 牧场与样方尺度下放牧强度测量及其对青藏高原可持续畜牧业发展的影响[J]. 生态学报, 2025, 45(10):4952-4962 LI Q, GAO S G, ZHANG C L, et al. Assessment of the impacts of climate change and human activities on the dynamic grassland change in Inner Mongolia[J]. Geography and Geo-Information Science, 2019, 35(3):99-104 李庆, 高素改, 张春来, 等. 内蒙古草地变化过程中气候变化和人类活动的相对作用评估[J]. 地理与地理信息科学, 2019, 35(3):99-104 WANG T T, ZHAO X L, LAN Y, et al. Rainfall interception characteristics of typical shrubs in the southwestern edge of maowusu desert[J]. Chinese Journal of Grassland, 2021, 43(5):82-89 王甜甜, 赵学琳, 兰艳, 等. 毛乌素沙地西南缘典型灌丛降雨截留特征[J]. 中国草地学报, 2021, 43(5):82-89 HOU H Y, ZHOU J W, LIU H J, et al. Advances in the study of shrub-herbs interaction of shrub encroachment in grassland[J]. Chinese Journal of Grassland, 2024, 46(5):132-141 侯慧扬, 周建伟, 刘慧娟, 等. 草原灌丛化过程灌木-草本植物相互作用研究进展[J]. 中国草地学报, 2024, 46(5):132-141 ZHAO K, ZHU Y, HA J, et al. Effects of grazing intensity and topography on carbon exchange in desert grassland[J]. Chinese Journal of Grassland, 2021, 43(12):40-46, 65 赵坤, 朱毅, 哈洁, 等. 放牧强度和地形对荒漠草原碳交换量的影响[J]. 中国草地学报, 2021, 43(12):40-46, 65 SUN Q , JIA R , TIAN J , et al.Interplay of grazing intensity and topography on soil phosphorus dynamics in semiarid grasslands[J].Catena, 2025(255):109072 WU L J. Effects of grazing on phosphorus balance in a typical steppe of Inner Mongolia[D]. Hohhot:Inner Mongolia Agricultural University, 2020:25-26 乌力吉. 放牧对内蒙古典型草原生态系统磷平衡的影响[D]. 呼和浩特:内蒙古农业大学, 2020:25-26 HOFFMANN C, FUNK R, WIELAND R, et al. Effects of grazing and topography on dust flux and deposition in the Xilingele grassland, Inner Mongolia[J]. Journal of Arid Environments, 2008, 72(5):792-807 LI W H, ZHENG S X, BAI Y F. Effects of grazing intensity and topography on species abundance distribution in a typical steppe of Inner Mongolia[J]. Chinese Journal of Plant Ecology, 2014, 38(2):178-187 李文怀, 郑淑霞, 白永飞. 放牧强度和地形对内蒙古典型草原物种多度分布的影响[J]. 植物生态学报, 2014, 38(2):178-187 BI X, YU T Q, SHI M Y, et al. The influence of grazing intensity on the composition of plant communities and ecological niches in the Leymus chinensis meadow steppe[J]. Acta Agrestia Scinca, 2026, 34(2):403-414 毕鑫, 俞天琦, 侍梦圆, 等. 放牧强度对羊草草甸草原植物群落组成及生态位的影响[J]. 草地学报, 2026, 34(2):403-414 GUO T T, GUO M Q, PANG Y, et al. Ungulate herbivores promote beta diversity and drive stochastic plant community assembly by selective defoliation and trampling: from a four-year simulation experiment[J]. Journal of Ecology, 2024, 112(9):1992-2006 JIN Y L, ZHU L, YAN Y Y, et al. Effect of slope position on phylogenetic diversity of plant community a case study in the xilingol typical steppe[J]. Arid Zone Research, 2018, 35(2):363-369 金一兰, 朱羚, 燕亚媛, 等. 坡位对群落系统发育多样性的影响——以锡林郭勒典型草原为例[J]. 干旱区研究, 2018, 35(2):363-369)
The Influence of Grazing and Topography on the Main Populations and Population-Community Relationship of Alpine Shrublands in the Qilian Mountains
PENG Bai-qing1, SANG Ya-zhuan1, NING Jiao1, HOU Fu-jiang1
Affiliations
1 China-Kazakhstan Belt and Road Joint Laboratory on Grassland Ecological Restoration, Key Laboratory of Grassland Livestock Industry Innovation, Ministry of Agriculture and Rural Affairs, Engineering Technology Research Center for Ecological Restoration and Utilization of Degraded Grassland in Northwest China, National Forestry and Grassland Administration, College of Grassland Agricultural Science and Technology, Lanzhou University, Lanzhou, Gansu Province 730020, China
Grazing and topography are key factors influencing the structure and function of alpine shrub ecosystems, but their effects on shrub populations and communities remain unclear. For this purpose, this study explored the effects of topography (three types of slope positions) and grazing (three distances from livestock enclosure) on the dominant species of alpine shrubland Caragana jubata and the companion species Thalictrum alpinum in the Qilian Mountain Grassland Agricultural Demonstration Area. The results showed that: 1) The distance from livestock enclosure explained the plant height and height-to-crown ratio of the Caragana jubata as 45.10% and 46.80% (P<0.01), and the plant height, height-to-crown ratio and density of the Thalictrum alpinum as 46.50%, 40.60%(P<0.01) and 55.30% (P<0.001). The topography only affects the population coverage of the Caragana jubata, which can explain that the coverage of the Caragana jubata is 22.30% (P<0.05). 2) The overlap degree of the ecological niches of the Caragana jubata and the Thalictrum alpinum is the lowest at a distance of 400 meters between the livestock enclosure and the highest at 1200 meters. 3) The plant height, crown breadth and density of the population of Thalictrum alpinum significantly affected the community coverage, with an interpretation rate of 35.3% (P<0.05). The community coverage significantly affected the population density of Thalictrum alpinum, with an interpretation rate of 25.1%(P<0.01). The plant height, density and importance value of the population of Caragana jubata significantly affected the community coverage, with an interpretation rate of 57.6%(P<0.01). This study reveals the role of grazing and topography in influencing the shrub-herbaceous competitive balance in alpine shrublands by regulating population growth and its population-community relationship, providing a scientific basis for grazing in complex topography.
Key words
Population
/
Community
/
Height
/
Crown breadth
/
Companion species
PENG Bai-qing, SANG Ya-zhuan, NING Jiao, HOU Fu-jiang.
The Influence of Grazing and Topography on the Main Populations and Population-Community Relationship of Alpine Shrublands in the Qilian Mountains[J].
Acta Agrestia Sinica,
2026
, 34
(5)
: 1827
-1840
.
DOI: 10.11733/j.issn.1007-0435.2026.05.024
HOU F J, YANG Z Y. Effects of grazing of livestock on grassland[J]. Acta Ecologica Sinica, 2006, 26(1):244-264 侯扶江, 杨中艺. 放牧对草地的作用[J]. 生态学报, 2006, 26(1):244-264 SONG W J, SU J S, ZHANG M D, et al. Plant compensatory growth and optimal grazing intensity of grasslands in northern China: A meta-analysis of grazing experiments[J]. Chinese Science Bulletin, 2023, 68(11):1330-1342 宋伟江, 苏纪帅, 张梦迪, 等. 中国北方草地植物补偿性生长与合理放牧强度:基于放牧实验的整合分析[J]. 科学通报, 2023, 68(11):1330-1342 JAMES M B, JOE F, MARK J S, et al.A plant trait analysis of responses to grazing in a long‐term experiment[J].Journal of Applied ecology, 2001, 38(2):253-267 FEI X, SUO C X, XIANG S, et al. Responses of plant community structure and functional group characteristics to long-term seasonal grazing in an alpine meadow on eastern Qinghai-Tibet Plateau[J]. Acta Agrestia Sinica, 2022, 30(8):1954-1963 费璇, 锁才序, 向双, 等. 青藏东缘高寒草甸植物群落结构及功能群特征对长期季节放牧的响应[J]. 草地学报, 2022, 30(8):1954-1963 DUAN H C, QI Y, KANG W P, et al. Seasonal variation of vegetation and its spatiotemporal response to climatic factors in the Qilian Mountains, China[J]. Sustainability, 2022, 14(9):4926 JING Y Y, XU C L, YU X J. The effect of grazing yak’s activity range on vegetation community structure and soil physicochemical characteristics in cold season pasture of alpine meadow[J]. Grassland and Turf, 2024, 44(4):54-63 景媛媛, 徐长林, 鱼小军. 牦牛放牧活动范围对高寒草甸冷季牧场植被群落结构和土壤理化性质的影响[J]. 草原与草坪, 2024, 44(4):54-63 BIAN J X, NIU X X, CHEN J, et al. Nest circle distance affects plant community structure and spatial distribution of soil nutrients in alpine meadows[J]. Pratacultural Science, 2024, 41(9):2003-2011 边金霞, 牛小霞, 陈娟, 等. 窝圈距离影响高寒草甸植物群落结构和土壤养分空间分布[J]. 草业科学, 2024, 41(9):2003-2011 SUN H, ZHAO Y G, LIU G L, et al. Effects of grazing intensity on the grassland vegetation community characteristics in the hilly Loess Plateau Region[J]. Acta Agrestia Sinica, 2022, 30(4):810-817 孙会, 赵允格, 刘广亮, 等. 放牧强度对黄土丘陵区草地植物群落特征的影响[J]. 草地学报, 2022, 30(4):810-817 WANG R H, PENG Q, ZHANG W D, et al. Ecohydrological service characteristics of Qilian Mountain ecosystem in the next 30 years based on scenario simulation[J]. Sustainability, 2022, 14(3):1819 ZHAO L P, BAI X, TAN S T, et al. Effect of different enclosure durations on aboveground vegetation in typical steppe on Loess Plateau[J]. Pratacultural Science, 2018, 35(1):27-35 赵凌平, 白欣, 谭世图, 等. 不同年限封育对黄土高原典型草原地上植被的影响[J]. 草业科学, 2018, 35(1):27-35 WANG L, XU D M, ZHANG J J. Effects of enclosure on composition of plant community and species diversity of desert steppe[J]. Pratacultural Science, 2012, 29(10):1512-1516 王蕾, 许冬梅, 张晶晶. 封育对荒漠草原植物群落组成和物种多样性的影响[J]. 草业科学, 2012, 29(10):1512-1516 PAN X X, LIU B R, NIU S F, et al. Effects of enclosure years on community structure and species diversity of desert grassland[J]. Guizhou Agricultural Sciences, 2018(7):95-99 潘笑笑, 刘秉儒, 牛宋芳, 等.围封年限对荒漠草原群落结构及物种多样性的影响[J].贵州农业科学, 2018, 46(7):95-99 XU H W, WANG X K, QU Q, et al. Variations and factors characterizing ecological niches of species in a stable grassland plant community[J]. Ecological Indicators, 2021, 128:107846 ZHANG F , BENNETT J A , ZHANG B , et al.Cessation of grazing stabilizes productivity through effects on species asynchrony and stability of shrub/semi-shrub plants in arid grasslands[J].Agriculture, Ecosystems & Environment, 2023(348):108411 WANG W , KANG Y , LIU M , et al.Shrub encroachment in alpine meadows facilitates soil carbon and nitrogen storage by redistributing soil water[J].Catena, 2025(256):109061 WANG L, MOU P, WANG T M. Impacts of topography on spatial variability of temperate understory biomass in closed forest: a case study of Northeast China Tiger and Leopard National Park[J]. Guihaia, 2024, 44(8):1512-1523 王乐, 牟溥, 王天明. 地形对温带密林下灌草层地上生物量分异的影响——以东北虎豹国家公园为例[J]. 广西植物, 2024, 44(8):1512-1523 LÜ Z L, LIU B, CHANG F, et al. Relationship between plant functional diversity and ecosystem multifunctionality in Bayanbulak alpine meadow along an altitude gradient[J]. Chinese Journal of Plant Ecology, 2023, 47(6):822-832 吕自立, 刘彬, 常凤, 等. 巴音布鲁克高寒草甸植物功能多样性与生态系统多功能性关系沿海拔梯度的变化[J]. 植物生态学报, 2023, 47(6):822-832 LI Q, RONG L, WANG M. Effects of topography on diversity and distribution pattern of plant species in karst mountains area[J]. Bulletin of Soil and Water Conservation, 2019, 39(6):27-34 李芹, 容丽, 王敏. 地形对喀斯特山地植物物种多样性及分布格局的影响[J]. 水土保持通报, 2019, 39(6):27-34 MA Y R, GUAN Q Y, SUN Y F, et al. Three-dimensional dynamic characteristics of vegetation and its response to climatic factors in the Qilian Mountains[J]. Catena, 2022, 208:105694 FENG X X, CHEN Q. Drivers grass-shrub competition in grassland ecosystems[J]. Chinese Journal of Grassland, 2024, 46(8):137-148 冯星星, 陈清. 草原生态系统影响草本-灌木竞争的主要因素[J]. 中国草地学报, 2024, 46(8):137-148 ZHANG P J, HUANG J H, MU L, et al. Influence of nitrogen and water addition on the primary productivity of Stipa breviflora in a desert steppe under different grazing intensities[J]. Acta Ecologica Sinica, 2022, 42(13):5458-5470 张璞进, 黄建辉, 木兰, 等. 氮水添加对放牧背景下荒漠草原生产力的影响[J]. 生态学报, 2022, 42(13):5458-5470 WANG J F, ZHANG L H, ZHAO R F, et al. Responses of plant growth of different life-forms to precipitation changes in desert steppe[J]. Chinese Journal of Applied Ecology, 2020, 31(3):778-786 王军锋, 张丽华, 赵锐锋, 等. 荒漠草原区不同生活型植物生长对降水变化的响应[J]. 应用生态学报, 2020, 31(3):778-786 GAO L X, LAN T Y, ZHAO Z X, et al. Spatial turnover of shrubland communities and underlying factors in northern mid-subtropical China[J]. Chinese Journal of Plant Ecology, 2022, 46(11):1411-1421 高璐鑫, 兰天元, 赵志霞, 等. 中国中亚热带北部灌丛群落植物空间周转及其驱动因素[J]. 植物生态学报, 2022, 46(11):1411-1421 WANG B B, ZHOU S Q, JING Y D, et al. Temporal-spatial variations of vegetation phenology and the influence of topography on phenology in Shanxi Province[J]. Chinese Journal of Ecology, 2021, 40(6):1839-1848 王贝贝, 周淑琴, 荆耀栋, 等. 山西省植被物候时空变化以及地形对物候的影响[J]. 生态学杂志, 2021, 40(6):1839-1848 XIAO X, GE R, HOU F J. Effects of grazing and topography on α diversity and physical properties of the soil of alpine meadow communities in the Qinghai-Tibet Plateau[J]. Pratacultural Science, 2019, 36(12):3041-3051 肖翔, 格日才旦, 侯扶江. 青藏高原放牧和地形对高寒草甸群落α多样性和土壤物理性质的影响[J]. 草业科学, 2019, 36(12):3041-3051 LI Y, ZHAO X R, LI G T, et al. Effect of grazing on soil microbial biomass phosphorus and phosphatase activities on different topographic unit of a typical steppe[J]. Acta Ecologica Sinica, 2022, 42(10):4137-4149 李营, 赵小蓉, 李贵桐, 等. 内蒙古典型草原不同地形单元放牧对土壤微生物量磷及磷酸酶活性的影响[J]. 生态学报, 2022, 42(10):4137-4149 HOWISON R A, OLFF H, VAN DE KOPPEL J, et al. Biotically driven vegetation mosaics in grazing ecosystems: the battle between bioturbation and biocompaction[J]. Ecological Monographs, 2017, 87(3):363-378 HOU S J, WANG Y X. Summer grazing of red deer effects on plant diversity of alpine grassland on Qilian Mountain[J]. Acta Prataculturae Sinica, 2020, 29(10):206-210 侯帅君, 王迎新. 马鹿夏季放牧对祁连山草原群落多样性的影响[J]. 草业学报, 2020, 29(10):206-210 REN J Z. Grassland Agroecology[M]. Beijing: China Agriculture Press, 1995: 51-84 任继周. 草地农业生态学[M]. 北京:中国农业出版社, 1995:51-84 LIU J X, GONG Y B, ZUO Q, et al.A review of mountain grazing-paths and vegetationspatial heterogeneity [J]. Acta Prataculturae Sinica, 2012, 21(1):254-261 刘金鑫, 宫渊波, 左琴, 等. 山地牧道与植被空间异质性研究评述[J]. 草业学报, 2012, 21(1):254-261 LU W D, ZHU H B. SPSS Statistical Analysis[M]. 5th ed. Beijing: Publishing House of Electronics Industry, 2015:286-301 卢纹岱, 朱红兵. SPSS统计分析[M]. 5版. 北京:电子工业出版社, 2015:286-301 SHIPLEY B. The AIC model selection method applied to path analytic models compared using a d-separation test[J]. Ecology, 2013, 94(3):560-564 ZHANG R, YU F Y, ZHOU R H, et al. Effects of slope position and aspect on structure and species diversity of shrub community in the Jiajin Mountains, Sichuan Province, China[J]. Chinese Journal of Applied Ecology, 2020, 31(8):2507-2514 张荣, 余飞燕, 周润惠, 等. 坡向和坡位对四川夹金山灌丛群落结构与物种多样性特征的影响[J]. 应用生态学报, 2020, 31(8):2507-2514 WANG Z H, LYU S J, WANG Z W, et al. Correlation analysis between the important value of dominant population and species diversity after spring rest grazing in desert steppe[J]. Chinese Journal of Grassland, 2025, 47(1):45-53 王梓晗, 吕世杰, 王忠武, 等. 荒漠草原春季休牧后优势种群重要值与物种多样性的关联性分析[J]. 中国草地学报, 2025, 47(1):45-53 QIN H, LI M G, GAO J, et al. Composition and diversity of soil bacterial communities in shrub at different altitudes in luya mountain[J]. Ecology and Environmental Sciences, 2023, 32(3):459-468 秦浩, 李蒙爱, 高劲, 等. 芦芽山不同海拔灌丛土壤细菌群落组成和多样性研究[J]. 生态环境学报, 2023, 32(3):459-468 FAN Y , LI X Y , HUANG H , et al.Does phenology play a role in the feedbacks underlying shrub encroachment?[J].Science of The Total Environment, 2018, 657(20):1064-1073 YANG W , QU G , KELLY A R , et al.Positive effects of leguminous shrub encroachment on multiple ecosystem functions of alpine meadows and steppes greatly depended on increasing soil nutrient[J].Catena, 2024(236):107745 LI Y F, CHEN L, LI X B, et al. Structural characteristics and ecological effects of a shrub-fertile island in a desert steppe[J]. Pratacultural Science, 2018, 35(10):2327-2335 李月飞, 陈林, 李学斌, 等. 荒漠草原灌丛沃岛的结构特征和生态效应[J]. 草业科学, 2018, 35(10):2327-2335 LIU Y Y, DING J Y. Research progress on shrub encroachment in China[J]. Chinese Journal of Grassland, 2025, 47(4):127-141 刘奕吟, 丁婧祎. 中国草地灌丛化研究进展[J]. 中国草地学报, 2025, 47(4):127-141 ZHANG X, YE M, CHEN W L, et al. Influence of topography on species diversity of grassland plant communities in the Altai Mountains[J]. Chinese Journal of Grassland, 2024, 46(7):57-71 张西, 叶茂, 陈维龙, 等. 地形对阿尔泰山草地植物群落物种多样性的影响[J]. 中国草地学报, 2024, 46(7):57-71 AKASH TARIQ, GAOY J, ZENG F J, et al. Guardians of arid lands: deep-rooted defense against desertification and climate change[J].Trends in Plant Science, 2026, 31(3):295-312 QIAO O M, CHEN Z. Plant diversity on different types of slope ecological engineering and its responses to environmental factors in mining areas[J]. Chinese Journal of Applied Ecology, 2022, 33(3):742-748 乔欧盟, 陈璋. 矿区不同类型生态护坡工程植物多样性对环境因子的响应[J]. 应用生态学报, 2022, 33(3):742-748 WANG W H, WANG S L, LIANG G L, et al. Effects of slope categories of differing aspect and position on plant community diversity in alpine shrubland in the Qilian Mountains[J]. Acta Prataculturae Sinica, 2025, 34(1):17-28 王文虎, 王世林, 梁国玲, 等. 坡向和坡位对祁连山高寒灌丛植物群落多样性的影响[J]. 草业学报, 2025, 34(1):17-28 SUN Y, HE M Z, WANG L. Effects of precipitation control on plant diversity and biomass in a desert region[J]. Acta Ecologica Sinica, 2018, 38(7):2425-2433 孙岩, 何明珠, 王立. 降水控制对荒漠植物群落物种多样性和生物量的影响[J]. 生态学报, 2018, 38(7):2425-2433 XIAO Y M, YANG L C, NIE X Q, et al. Effects of slope position on phylogenetic diversity and structure of alpine shrub community[J]. Chinese Journal of Ecology, 2019, 38(6):1611-1619 肖元明, 杨路存, 聂秀青, 等. 坡位对高寒灌丛群落谱系多样性和谱系结构的影响[J]. 生态学杂志, 2019, 38(6):1611-1619 CAO J Y, LIU J F, YUAN Q, et al. Traits of shrubs in forests and bushes reveal different life strategies[J]. Chinese Journal of Plant Ecology, 2020, 44(7):715-729 曹嘉瑜, 刘建峰, 袁泉, 等. 森林与灌丛的灌木性状揭示不同的生活策略[J]. 植物生态学报, 2020, 44(7):715-729 LIU Y Z, SUN C C, LIU W T, et al. Response of keystone species changes in alpine grassland plant communities to different herbivore assemblage grazing[J]. Acta Ecologica Sinica, 2022, 42(18):7529-7540 刘玉祯, 孙彩彩, 刘文亭, 等. 高寒草地植物群落关键种对不同放牧家畜组合放牧的响应[J]. 生态学报, 2022, 42(18):7529-7540 JIANG K W, ZHANG Q Q, WANG Y F, et al. Characteristics of plant functional groups and the relationships with soil environmental factors in middle part of northern slope of Tianshan Mountains under different grazing intensities[J]. Chinese Journal of Plant Ecology, 2024, 48(6):701-718 江康威, 张青青, 王亚菲, 等. 放牧干扰下天山北坡中段植物功能群特征及其与土壤环境因子的关系[J]. 植物生态学报, 2024, 48(6):701-718 HAN D Y, YANG Y F, ZHANG W, et al. Impact of cutting time and intensity simulating large herbivore grazing on compensatory growth of Elymus dahuricus[J]. Acta Ecologica Sinica, 2026, 46(1):582-592 韩大勇, 杨允菲, 张维, 等. 刈割强度与频率模拟大型草食动物采食对披碱草补偿生长的影响[J]. 生态学报, 2026, 46(1):582-592 ZHANG Y F, YANG X X, DONG Q M, et al. Effects of Mixed Grazing of Yak and Tibetan Sheep on Feed Intake of Grazing Livestock and Plant Compensation Growth[J]. Acta Agrestia Sinica, 2019, 27(6):1607-1614 张艳芬, 杨晓霞, 董全民, 等.牦牛和藏羊混合放牧对放牧家畜采食量和植物补偿性生长的影响[J].草地学报, 2019, 27(6):1607-1614 YANG Y, LIU A J, LI L H, et al. Effects of different disturbance types on plant species composition and functional group characteristics of typical steppe in Inner Mongolia, China[J]. Chinese Journal of Applied Ecology, 2016, 27(3):794-802 杨勇, 刘爱军, 李兰花, 等. 不同干扰方式对内蒙古典型草原植物种组成和功能群特征的影响[J]. 应用生态学报, 2016, 27(3):794-802 CANALS R M, MÚGICA L, DURÁN M, et al. Restorative pyric herbivory practices in shrub-encroached grasslands enhance nutrient resource availability and spatial heterogeneity[J]. Agriculture, Ecosystems & Environment, 2024(372):109072 MI Y Q, XU H B, ZHANG L, et al. The impact of enclosure on vegetation ecological niche and interspecific linkage in Inner Mongolian desert steppe[J]. Acta Agrestia Sinica2026, 34(3):1007-1016 米雨晴, 许宏斌, 张雷, 等. 围封对内蒙古荒漠草原植被生态位及种间联结性的影响[J]. 草地学报, 2026, 34(3):1007-1016 LI L, LI D, MHKNOPS J, et al. Assessing grazing intensity at pasture and quadrat scales for sustainable pastoralism on the Qinghai-Tibetan Plateau[J]. Acta Ecologica Sinica, 2025, 45(10):4952-4962 李黎, 李丹, MHKNOPS Johannes, 等. 牧场与样方尺度下放牧强度测量及其对青藏高原可持续畜牧业发展的影响[J]. 生态学报, 2025, 45(10):4952-4962 LI Q, GAO S G, ZHANG C L, et al. Assessment of the impacts of climate change and human activities on the dynamic grassland change in Inner Mongolia[J]. Geography and Geo-Information Science, 2019, 35(3):99-104 李庆, 高素改, 张春来, 等. 内蒙古草地变化过程中气候变化和人类活动的相对作用评估[J]. 地理与地理信息科学, 2019, 35(3):99-104 WANG T T, ZHAO X L, LAN Y, et al. Rainfall interception characteristics of typical shrubs in the southwestern edge of maowusu desert[J]. Chinese Journal of Grassland, 2021, 43(5):82-89 王甜甜, 赵学琳, 兰艳, 等. 毛乌素沙地西南缘典型灌丛降雨截留特征[J]. 中国草地学报, 2021, 43(5):82-89 HOU H Y, ZHOU J W, LIU H J, et al. Advances in the study of shrub-herbs interaction of shrub encroachment in grassland[J]. Chinese Journal of Grassland, 2024, 46(5):132-141 侯慧扬, 周建伟, 刘慧娟, 等. 草原灌丛化过程灌木-草本植物相互作用研究进展[J]. 中国草地学报, 2024, 46(5):132-141 ZHAO K, ZHU Y, HA J, et al. Effects of grazing intensity and topography on carbon exchange in desert grassland[J]. Chinese Journal of Grassland, 2021, 43(12):40-46, 65 赵坤, 朱毅, 哈洁, 等. 放牧强度和地形对荒漠草原碳交换量的影响[J]. 中国草地学报, 2021, 43(12):40-46, 65 SUN Q , JIA R , TIAN J , et al.Interplay of grazing intensity and topography on soil phosphorus dynamics in semiarid grasslands[J].Catena, 2025(255):109072 WU L J. Effects of grazing on phosphorus balance in a typical steppe of Inner Mongolia[D]. Hohhot:Inner Mongolia Agricultural University, 2020:25-26 乌力吉. 放牧对内蒙古典型草原生态系统磷平衡的影响[D]. 呼和浩特:内蒙古农业大学, 2020:25-26 HOFFMANN C, FUNK R, WIELAND R, et al. Effects of grazing and topography on dust flux and deposition in the Xilingele grassland, Inner Mongolia[J]. Journal of Arid Environments, 2008, 72(5):792-807 LI W H, ZHENG S X, BAI Y F. Effects of grazing intensity and topography on species abundance distribution in a typical steppe of Inner Mongolia[J]. Chinese Journal of Plant Ecology, 2014, 38(2):178-187 李文怀, 郑淑霞, 白永飞. 放牧强度和地形对内蒙古典型草原物种多度分布的影响[J]. 植物生态学报, 2014, 38(2):178-187 BI X, YU T Q, SHI M Y, et al. The influence of grazing intensity on the composition of plant communities and ecological niches in the Leymus chinensis meadow steppe[J]. Acta Agrestia Scinca, 2026, 34(2):403-414 毕鑫, 俞天琦, 侍梦圆, 等. 放牧强度对羊草草甸草原植物群落组成及生态位的影响[J]. 草地学报, 2026, 34(2):403-414 GUO T T, GUO M Q, PANG Y, et al. Ungulate herbivores promote beta diversity and drive stochastic plant community assembly by selective defoliation and trampling: from a four-year simulation experiment[J]. Journal of Ecology, 2024, 112(9):1992-2006 JIN Y L, ZHU L, YAN Y Y, et al. Effect of slope position on phylogenetic diversity of plant community a case study in the xilingol typical steppe[J]. Arid Zone Research, 2018, 35(2):363-369 金一兰, 朱羚, 燕亚媛, 等. 坡位对群落系统发育多样性的影响——以锡林郭勒典型草原为例[J]. 干旱区研究, 2018, 35(2):363-369